Processing of file "Santanachelys*" begins...
Data matrix has 18 taxa, 104 characters
Valid character-state symbols: 0123
"Equate" macros in effect:
a ==> {01}
b ==> {12}
c ==> {02}
d ==> {012}
e ==> {123}
Missing data identified by '?'
Processing of file "Santanachelys*" completed.
Branch-and-bound search settings:
Initial upper bound: unknown (compute via stepwise)
Addition sequence: furthest
Initial MAXTREES setting = 100
Branches having maximum length zero collapsed to yield polytomies
Topological constraints not enforced
Trees are unrooted
Multi-state taxa interpreted as uncertainty
Branch-and-bound search completed:
Shortest tree found = 208
Number of trees retained = 1
Time used = 00:43:04.7
Tree description:
Unrooted tree(s) rooted using outgroup method
Character-state optimization: Delayed transformation (DELTRAN)
Tree number 1:
Tree length = 208
Consistency index (CI) = 0.567
Homoplasy index (HI) = 0.433
CI excluding uninformative characters = 0.554
HI excluding uninformative characters = 0.446
Retention index (RI) = 0.720
Rescaled consistency index (RC) = 0.409
/------Plesiochelyidae
| /------Xinjiangchelys
| | /------Sinemydidae
\---34 | /------Chelydridae
\----33 | /------Chelomacryptodir
\---32 | /------Toxochelys
| | /------22 /------Ctenochelys
\---31 | \---21 /------Osteopygis
| | \---20 /------Puppigerus
| | \----19------Chelonia
\---30 /------Corsochelys
| /------28 /------Mesodermochelys
| | \----27------Dermochelys
\----29 /------Santanachelys
\---26 /------Notochelone
\---25 /------Desmatochelys
\---24 /------Chelosphargis
\----23------Protostega gigas
Apomorphy lists:
Branch Character Steps CI Change
Plesiochelyidae <-> node_34 8.posterior 1 0.400 01 <=> 2
51.8th cervic 1 1.000 0 <=> 1
58.anterior p 1 1.000 0 <=> 1
59.posterior 1 1.000 0 <=> 1
92.axillary b 1 1.000 0 <=> 1
93.inguinal b 1 1.000 0 <=> 1
94.femonal an 1 0.500 0 <=> 1
node_34 --> Xinjiangchelys 95.plastral i 1 0.400 0 --> 1
node_34 --> node_33 11.jugal pter 1 0.500 1 --> 0
20.palatines 1 0.500 1 --> 0
33.dorsum sel 1 0.500 1 --> 0
43.lingual ri 1 0.333 0 ==> 1
45.splenial 1 1.000 0 --> 1
49.cervical c 1 0.667 0 ==> 1
50.biconvex a 1 0.667 0 ==> 2
95.plastral i 1 0.400 0 --> 2
98.narrow epi 1 1.000 0 ==> 1
102.narrow xip 1 1.000 0 ==> 1
node_33 --> Sinemydidae 1.cranial sc 1 0.200 1 --> 0
5.orbital di 1 0.333 0 --> 1
21.foramen pa 1 0.500 0 --> 2
35.short rost 1 0.500 0 ==> 1
46.transverse 1 1.000 1 ==> 0
56.10th thora 1 0.500 0 --> 1
83.1st vertbr 1 0.200 0 --> 1
node_33 --> node_32 2.nasal 0. 1 0.250 0 --> 1
30.internal c 1 0.667 1 ==> 2
36.split pala 1 1.000 0 ==> 1
47.proportion 1 0.333 0 ==> 1
52.8th cervic 1 0.333 0 ==> 1
54.1st thorac 1 0.500 0 ==> 1
97.plastral f 1 0.667 0 --> 1
node_32 --> Chelydridae 4.prefrontal 1 0.250 0 ==> 1
13.hooked pre 1 0.500 0 ==> 1
40.hooked den 1 0.500 0 ==> 1
48.ventral th 1 0.500 0 --> 1
53.double cer 1 0.333 0 --> 1
node_32 --> node_31 57.chevron 1 1.000 0 ==> 1
59.posterior 1 1.000 1 ==> 2
node_31 --> Chelomacryptodir 53.double cer 1 0.333 0 --> 1
56.10th thora 1 0.500 0 --> 1
94.femonal an 1 0.500 1 ==> 0
node_31 --> node_30 6.processus 1 1.000 1 ==> 0
8.posterior 1 0.400 2 --> 0
14.foramen pr 1 1.000 0 ==> 1
29.paired for 1 1.000 0 ==> 1
33.dorsum sel 1 0.500 0 ==> 1
41.symphyseal 1 0.250 0 --> 1
55.1st thorac 1 0.500 0 --> 1
60.coracoid 1 1.000 0 --> 1
66.humerus 1 0.500 0 --> 1
67.lateral pr 1 0.667 0 --> 1
76.flat carpa 1 1.000 0 ==> 1
77.3rd to 5th 1 1.000 0 ==> 1
89.pleural re 1 0.400 0 ==> 1
91.plastral s 1 1.000 0 --> 1
99.entoplastr 1 0.500 0 ==> 1
node_30 --> node_22 5.orbital di 1 0.333 0 --> 1
7.parietal s 1 0.333 1 ==> 0
15.upper trit 1 1.000 0 ==> 1
18.vomer pala 1 1.000 0 ==> 1
31.V shaped c 1 1.000 0 ==> 1
48.ventral th 1 0.500 0 --> 1
74.ulna radiu 1 1.000 0 ==> 1
82.cervical s 1 0.500 0 --> 1
85.neural num 1 0.500 0 ==> 1
node_22 --> Toxochelys 1.cranial sc 1 0.200 1 --> 0
2.nasal 0. 1 0.250 1 ==> 0
8.posterior 1 0.400 0 ==> 1
21.foramen pa 1 0.500 0 --> 2
81.posterior 1 0.500 0 --> 1
node_22 --> node_21 16.upper trit 1 0.333 0 --> 1
23.median ven 1 1.000 0 ==> 1
32.V shaped b 1 1.000 0 ==> 1
34.rod like r 1 0.500 0 ==> 1
node_21 --> Ctenochelys 1.cranial sc 1 0.200 1 --> 0
52.8th cervic 1 0.333 1 ==> 0
81.posterior 1 0.500 0 --> 1
84.thick neur 1 0.333 0 ==> 1
86.neural sha 1 1.000 0 ==> 1
node_21 --> node_20 12.lower chee 1 0.500 1 ==> 0
17.vomerine p 1 1.000 0 ==> 1
21.foramen pa 1 0.500 0 --> 3
37.foramen ca 1 0.500 0 ==> 1
53.double cer 1 0.333 0 ==> 1
node_20 --> Osteopygis 39.broad lowe 1 0.500 0 --> 1
41.symphyseal 1 0.250 1 ==> 0
66.humerus 1 0.500 1 ==> 0
67.lateral pr 1 0.667 1 ==> 0
83.1st vertbr 1 0.200 0 --> 1
89.pleural re 1 0.400 1 --> 0
99.entoplastr 1 0.500 1 ==> 0
node_20 --> node_19 5.orbital di 1 0.333 1 ==> 0
49.cervical c 1 0.667 1 ==> 2
64.lateral pr 1 0.500 0 ==> 1
68.shoulder o 1 0.500 0 ==> 1
70.V shaped l 1 1.000 0 ==> 1
78.1st and 2n 1 0.333 0 ==> 1
79.femoral tr 1 0.500 0 ==> 1
node_19 --> Puppigerus 39.broad lowe 1 0.500 0 --> 1
89.pleural re 1 0.400 1 --> 0
95.plastral i 1 0.400 2 --> 1
node_19 --> Chelonia 38.foramen ca 1 0.500 0 ==> 1
42.high symph 1 0.500 0 ==> 1
43.lingual ri 1 0.333 1 ==> 0
44.anterior s 1 0.500 0 ==> 1
61.scapular a 1 0.500 0 ==> 1
69.scar for M 1 0.333 0 ==> 1
83.1st vertbr 1 0.200 0 --> 1
95.plastral i 1 0.400 2 --> 0
103.medial con 1 0.333 0 ==> 1
node_30 --> node_29 10.medial pro 1 1.000 0 ==> 1
11.jugal pter 1 0.500 0 ==> 1
47.proportion 1 0.333 1 ==> 0
52.8th cervic 1 0.333 1 ==> 0
54.1st thorac 1 0.500 1 ==> 0
61.scapular a 1 0.500 0 ==> 1
63.thyroid fe 1 1.000 0 ==> 1
64.lateral pr 1 0.500 0 ==> 1
68.shoulder o 1 0.500 0 ==> 1
79.femoral tr 1 0.500 0 ==> 1
86.neural sha 1 1.000 0 ==> 2
95.plastral i 1 0.400 2 --> 0
101.entoplastr 1 1.000 0 ==> 1
node_29 --> node_26 2.nasal 0. 1 0.250 1 ==> 0
9.jugal quad 1 1.000 0 ==> 1
12.lower chee 1 0.500 1 ==> 0
20.palatines 1 0.500 0 ==> 1
21.foramen pa 1 0.500 0 --> 1
24.pterygoid 1 1.000 0 ==> 1
30.internal c 1 0.667 2 --> 0
34.rod like r 1 0.500 0 ==> 1
50.biconvex a 1 0.667 2 ==> 1
62.lateral pr 1 0.500 0 --> 1
71.expansion 1 1.000 0 ==> 1
75.radius cur 1 1.000 0 ==> 1
82.cervical s 1 0.500 0 --> 1
84.thick neur 1 0.333 0 ==> 1
node_26 --> Santanachelys 3.medial mee 1 0.333 1 --> 0
55.1st thorac 1 0.500 1 ==> 0
88.suprapygal 1 0.500 0 --> 1
node_26 --> node_25 16.upper trit 1 0.333 0 --> 1
19.high lingu 1 1.000 0 ==> 1
72.medial con 1 1.000 0 ==> 1
78.1st and 2n 1 0.333 0 --> 1
103.medial con 1 0.333 0 --> 1
104.short xiph 1 1.000 0 ==> 1
node_25 --> Notochelone 4.prefrontal 1 0.250 0 --> 1
node_25 --> node_24 21.foramen pa 1 0.500 1 --> 3
25.pterygoid 1 1.000 0 ==> 1
26.large proc 1 1.000 0 ==> 1
88.suprapygal 1 0.500 0 --> 1
node_24 --> Desmatochelys 3.medial mee 1 0.333 1 --> 0
8.posterior 1 0.400 0 ==> 1
83.1st vertbr 1 0.200 0 ==> 1
84.thick neur 1 0.333 1 ==> 0
97.plastral f 1 0.667 1 ==> 2
node_24 --> node_23 4.prefrontal 1 0.250 0 --> 1
69.scar for M 1 0.333 0 ==> 1
96.star shape 1 1.000 0 ==> 1
node_23 --> Chelosphargis 3.medial mee 1 0.333 1 --> 0
node_23 --> Protostega gigas 1.cranial sc 1 0.200 1 ==> 0
2.nasal 0. 1 0.250 0 ==> 1
7.parietal s 1 0.333 1 ==> 0
13.hooked pre 1 0.500 0 ==> 1
22.processus 1 0.500 0 ==> 1
41.symphyseal 1 0.250 1 ==> 0
65.large pelv 1 1.000 0 ==> 1
80.scute sulc 1 0.500 0 ==> 1
85.neural num 1 0.500 0 ==> 1
89.pleural re 1 0.400 1 ==> 2
90.9th pelura 1 1.000 0 ==> 1
100.epi and en 1 1.000 0 ==> 1
node_29 --> node_28 1.cranial sc 1 0.200 1 --> 0
27.processus 1 1.000 0 ==> 1
35.short rost 1 0.500 0 ==> 1
37.foramen ca 1 0.500 0 ==> 1
80.scute sulc 1 0.500 0 ==> 1
97.plastral f 1 0.667 1 ==> 2
103.medial con 1 0.333 0 --> 1
node_28 --> Corsochelys 8.posterior 1 0.400 0 ==> 2
49.cervical c 1 0.667 1 ==> 0
50.biconvex a 1 0.667 2 ==> 0
83.1st vertbr 1 0.200 0 --> 1
node_28 --> node_27 28.crista sup 1 1.000 1 ==> 0
44.anterior s 1 0.500 0 --> 1
62.lateral pr 1 0.500 0 --> 1
73.elongate l 1 1.000 0 ==> 1
78.1st and 2n 1 0.333 0 --> 1
node_27 --> Mesodermochelys 42.high symph 1 0.500 0 ==> 1
43.lingual ri 1 0.333 1 ==> 0
87.neural red 1 1.000 0 --> 1
88.suprapygal 1 0.500 0 --> 1
node_27 --> Dermochelys 4.prefrontal 1 0.250 0 --> 1
7.parietal s 1 0.333 1 --> 0
16.upper trit 1 0.333 0 --> 1
21.foramen pa 1 0.500 0 --> 3
22.processus 1 0.500 0 --> 1
30.internal c 1 0.667 2 --> 0
38.foramen ca 1 0.500 0 --> 1
40.hooked den 1 0.500 0 ==> 1
41.symphyseal 1 0.250 1 ==> 0
47.proportion 1 0.333 0 ==> 1
67.lateral pr 1 0.667 1 ==> 2
69.scar for M 1 0.333 0 ==> 1
87.neural red 1 1.000 0 --> 2
88.suprapygal 1 0.500 0 --> 2
89.pleural re 1 0.400 1 ==> 2