Atcw (1) ------MTKEVCSNIGLWLLLTLLIGNYV

Atvac (1) -MASTEALLPVTSLQDPLSESRSDQIPETRRRRPIKVHLAVYSGLLLIALYVTLIVTHDG

GU252170 (1) MEASSSTSHDPALFHAPLLYHP-----RRRSSRPLKGFAVIIGSVVFLLSLVTLIVNQSP

FJ864677 (1) MEASYDPEQNPP-LHAPLLDR------SSPRTRRFAAIFACLVSLLLLLALIINQAQ

Atcw (24) VNLEAS------HHVYKRLTQSTNTK------SPS------VNQPYR

Atvac (60) SKAEIATE------SRPR--MAGVSEKSNDGVWISSDDGKVEAFPWNNTILSWQR

GU252170 (56) EPLASNPSSVTEAGSYSMAAQPRGIAEGVSAKSNP------SLFDKVGFNWTNAMFYWQR

FJ864677 (51) QPFEK------VQPRGVAEGVSAKSNP------SLLNQVPFNWTNAMFSWQR

^ < ^

Atcw (53) TGFHFQPPKNWMNDPNGPMIYKGIYHLFYQWNPKGAVWGNIVWAHSTSTDLINWDPHPPA

Atvac (107) TAFHFQPEKNWMNDPNGPLFYKGWYHFFYQYNPNAAVWGDIVWGHAVSKDLIHWLYLPIA

GU252170 (110) TAYHFQPQKNWMNDPDGPLYHKGWYHLFYQYNPDSAIWGNITWGHAVSTDLIHWFYLPLA

FJ864677 (91) SAYHFQPQKNWMNDPNGPLYHKGWYHLFYQYNPYSAIWGNITWGHAVSRDLIHWLYLPLA

^

Atcw (113) IFPSAPFDINGCWSGSATILPNGKPVILYTGIDPKNQQVQNIAEPKNLSDPYLREWKKSP

Atvac (167) MVPDQWYDANGVWTGSATFLDDGSIVMLYTGSTDEFVQVQNLAYPEDPSDPLLLKWVKFS

GU252170 (170) MVPDQWYDINGCWTGSATLLPDGRIVMLYTGSTNDSVQVQNLAYPANLSDPLLLQWLKYP

FJ864677 (151) LVPDHWYDIKGVWTGSATILADGQIIMLYTGETNESVQVQNLAYPANVSDPLLLHWLKYP

<

Atcw (173) LNPLMAPDAVNGINASSFRDPTTAWLGQDKKWRVIIGSKIH-RRGLAITYTSKDFLKWEK

Atvac (227) GNPVLVP--PPGIGAKDFRDPTTAWKTSSGKWRITIGSKIN-RTGISLIYDTTDFKTYEK

GU252170 (230) GNPVVVP--PTGIEDEEFRDPTTAWLGPDGSWRIVVGTRFNTTIGTALVFQTTNFSDYEL

FJ864677 (211) GNPVMVP--PPGVKPDDFRDPTTAWLGPDGTWRLTMGSKFDTTIGISLVYHTTNFRDYEL

< < <

Atcw (232) SPEPLHYDDGSGMWECPDFFPVTRFGSNGVETSSFGEPNEILKHVLKISLDDTKHDYYTI

Atvac (284) HETLLHQVPNTGMWECVDFYPVSKTQLNGLDTSVNGPD---VKHVIKASMDDTRIDHYAI

GU252170 (288) LDGVLHAVPGTGMWECVDFYPVAINGSVGLDTTALGPG---IKHVLKASLDDTKVDHYAI

FJ864677 (269) LDGVLHAVPGTGMWECVDFYPVAINGSVALDTSSLGPG---IKHVLKASLDNTKVDHYAI

Atcw (292) GTYDRVKDKFVPDNGFKMDGTAP--RYDYGKYYASKTFFDSAKNRRILWGWTNESSSVED

Atvac (341) GTYDDSNATWVPDNPSIDVGISTGLRYDYGKYYASKTFYDQNKGRRILWGWIGESDSEAA

GU252170 (345) GTYDMITDKWTPDNPEEDVGIG--LKVDYGRYYASKTFFDQSKQRRILYGWVNETDSEAD

FJ864677 (326) GTYDPITDKWTPDNPEEDVGIG--LKVDYGRYYASKTFFDQHKQRRVLWGWINETDTETA

Atcw (350) DVEKGWSGIQTIPRKIWLDR-SGKQLIQWPVREVERLRTKQVKNLRNKVLKSGSRLEVYG

Atvac (401) DVQKGWSSVQGIPRTVVLDTRTHKNLVQWPVEEIKSLRLSSKK-FDMTIG-PGTVVPVDV

GU252170 (403) DLEKGWASIQTIPRSVLYDNKTGTHLLQWPVEEVESLRLNATV-FKDVVVEAGSVVPLDI

FJ864677 (384) DLKKGWASLQTIPRTVLYDNKTGTNLLQWPVEEVESLRLNSTM-FKEVLVEPGSVVPLDI

Atcw (409) VTAAQADVEVLFKVRDLEKADVIEPSWTDPQLICS---KMNVSVKSGLGPFGLMVLASKN

Atvac (459) GSATQLDIEAEFEIKTDDLKLFFDDDSVEADNKFSCETNGGSTARGALGPFGFSVLADEG

GU252170 (462) GTATQLDILAEFEIETLVLNSTEDEVSDCGD------GAVDRSTYGPFGVLVIADDS

FJ864677 (443) GTTTQLDILAEFEIEPLIP-STTNEIDNCGD------GAVDRSTYGPFGLLVIADAS

Atcw (466) LEEYTSVYFRIFKARQNSNKYVVLMCSDQSRSSLKEDNDKTTYGAFVDINPHQPLSLRAL

Atvac (519) LSEQTPVYFYVTKGKHS--KLNTVFCTDTSRSTLANDVVKPIYGSFVPVLKGEKLTMRIL

GU252170 (513) LSELTPIYFRPLNTSDG--SLETYFCADETRSSKAPDVTKRVYGGKIPVLDDENYNMRVL

FJ864677 (493) LSELTPIYFRPLNASDG--SLKTYFCSDETRSSKASDVFKQVYGGKVPVLDDENYNMRVL

Atcw (526) IDHSVVESFGGKGRACITSRVYPKLAIGKSSHLFAFNYGYQSVDVLNLNAWSMNSAQIS-

Atvac (577) VDHSIVEGFAQGGRSCITSRVYPTKAIYGATKLFLFNNAIDATVTASFTVWQMNNAFIHP

GU252170 (571) VDHSVVESFGGGGRTVITSRVYPTEAIYGAARLFLFNNASGVNVKATLKIWEMNSAFIRP

FJ864677 (551) VDHSIVESFAQGGRTVISSRIYPTEAIYGAARLFLFNNATGVNVKATLKIWELNSAFIRP

Atcw (585) ------

Atvac (637) YSSDDLGVPSST---

GU252170 (631) FPFEETLFQEMVAST

FJ864677 (611) FPFERDSV------

Supplementary data 1. Alignment of Arabidopsis cell-wall invertase, Arabidopsis vacuolar invertase with two cotton vacuolar invertases. Amino acids marked with < are involved in substrate binding and amino acids marked with a ^ are tryptophanes involved in the active site. The putative cleavage sites for the leader sequence are underlined.