Potential cell death cleavage sites highlighted in bovine PARP amino acid sequence.
A. Amino acid sequence of bovine PARP
ORIGIN
10 20 30 40 50 60
MAESSDKLYRVEYAKSGRAS CKKCKESIPKDSIRMAFMVE SPMFDGKIPH WYHLSCFWKV
70 80 90 100 110 120
GFSIWHPDVE VEGFSELRWD DQQTIKKMAE TGGRTDVSGK GQDGVGSKTE KTLIDFGAGY
130 140 150 160 170 180
AKSNRSTCKS CMEKIDKGQV RLSKKVVYPD KPQLGMVDCW YHPKCFVQKR EELGFRPEFS
190 200 210 220 230 240
ATHLMGFSVL TAEDQETLKK QLPAIKGERK RKGDEVDGIDEVTKKKSKKE KDKEIKLEKA
250 260 270 280 290 300
LKAQNDLIWN VKDELKKACS TNDLKELLIF NKQEVPSGES AILDRVADGMVFGALLPCEE
↓ 310 320 330 340 350 360
CSGQLVFKGDAYYCTGDVTA WTKCMVKTQT PNRKEWVTPK EFREISYFKKLKIKKQDRIF
370 380 390 400 410 420
PPESSTPVGA AAPPSAASAP AAVHSGPPDKPLSNMKILTL GKLSQNKDEV KATIEKLGGK
430 440 450 460 470 480
LTGTANKASL CISTKKEVDK LNKKMEEVKE ANIRVVSEDF LQDISASTKS LQELLSTHLL
490 500 510 520 530 540
SPWGAEVKVEPVEAVGPKGKSGAAPSKKSKGPVKEEGTNK SEKRMKLTLKGGAAVDPDSG
550 560 570 580 590 600
LEHNAHVLEK GGKVFSATLG LVDIVKGTNS YYKLQLLEDDKESRYWIFRSWGRVGTVIGS
610 620 630 640 650 660
NKLEQMPSKE DAIEHFMKLYEEKTGNAWHS KNFTKHPKKF YPLEIDYGQDEEAVKKLTVN
670 680 690 700 710 720
PGTKSKLPKPVQNLIKMIFD VESMKKAMVEYEIDLQKMPLGKLSKRQIQA AYSILSEVQQ
730 740 750 760 770 780
ALSQGSSDSH ILDLSNRFYT LIPHDFGMKKPPLLNNANSV QAKVEMLDNL LDIEVAYSLL
790 800 810 820 830 840
RGGSDDSSKD PIDVNYEKLK TDIKVVDKDSEEAEIIRKYV KNTHATTHNA YDLEVVDIFK
850 860 870 880 890 900
IEREGESQRY KPFKQLHNRRLLWHGSRTTN FAGILSQGLR IAPPEAPVTG YMFGKGIYFA
910 920 930 940 950 960
DMVSKSANYC HTSQGDPIGL ILLGEAALGN MYELKHARHI SKLPKGKHSV KGLGKTTPDP
970 980 990 1000 1010
SASITVDGVEVPLGTGISSG VNDTCLLYNEYIVYDIAQVH LKYLLKLKFN FKTSLW//
END.*Cleavage sites based on synthetic substrates and literature.
B.
Legend of highlighted potential cleavage sitesFR / cathepsin B & L
DEVD / caspase 3
VAD / partial caspase 2
S_QLV__G / putative N-terminus of our fragment
IXXD, DXD / granzyme B
LY / calpain class I and II
H384, L480, G502, T658 / calpain*
RR, T528, T558 / cathepsin B [25]
L527, K555 / cathepsin G [24]
LEXD / caspases -8, -9, 10
YEXD / caspases -1, -4, -5, 14
* Buki, K. G., Bauer, P. I., and Kun, E. (1997) Biochim, Biophys. Acta 1338:100-106.
* Tompa, P., Buzder-Lantos, P., Tantos, A., et al (2004) J. Biol. Chem. 279: 20775-20785.
Figure S1. Potential cell death cleavage sites highlighted in bovine PARP amino acid sequence.
Our ~50 kDa PARP fragment was sent to the W.M. Keck Biotechnology Resource Lab at Yale University, and the sequence was read by Nancy Williams. Results were compared to the amino acid sequence for bovine PARP obtained from the National Center for Biotechnology Information (NCBI) Sequence Viewer: =protein&val=130779. Potential cleavage sites for various enzymes are highlighted throughout the PARP sequence (A). Sites correspond to either commercially available synthetic substrates or cleavage sites mentioned in the literature (see legend, B). Amino acids matching the results of the first 5 cycles of Edman (N-terminal) sequencing are highlighted beginning at a.a. 302 (S*GLV**G). Hence, the putative N-terminus of our fragment does not correspond to any of the possible sites we have located within the sequence. The suggestion is that more than one protease, including perhaps an exopeptidase, is active during the peak period of degeneration.