Electronic Supplementary Material Figure Captions

Electronic Supplementary Material

Modeling invasion risk for coastal marine species utilizing environmental and transport vector data

Electronic Supplementary Material Figure Captions:

ESM Fig. 1 Projected binary environmental suitability (ES) for models trained with only Global Biodiversity Information Facility (GBIF) occurrence records and corresponding bias grids showing suitable environments based on minimum only (yellow) and minimum and 10 percent (red) training presence logistic threshold. Blue represents areas that are not modeled as suitable. A: Carcinus maenas; B: Charybdis hellerii; C: Charybdis japonica; D: Hemigrapsus sanguineus; E: Rhithropanopeus harrisii.

ESM Fig. 2 Projected binary environmental suitability (ES) for models trained with GBIF and literature occurrence records without the inclusion of a bias grid showing suitable environments based on minimum only (yellow) and minimum and 10 percent (red) training presence logistic thresholds. Blue represents areas that are not modeled as suitable. A: Carcinus maenas; B: Charybdis hellerii; C: Charybdis japonica; D: Hemigrapsus sanguineus; E: Rhithropanopeus harrisii.

ESM Fig. 3 Occurrence record locations (native and non-native from all sources) for Carcinus maenas, Charybdis hellerii, Charybdis japonica, Hemigrapsus sanguineus, and Rhithropanopeus harrisii along with locations of major world ports (N=208; black triangles).

Contents:

GBIF Citations and Acknowledgements

Supplementary Literature Citations

R Code

Please contact the author for information regarding more details concerning port lists, R code, and specific data used in this manuscript beyond what is included within.

GBIF Citations and acknowledgements

GBIF search term “Carcinus maenas”

ArtDatabanken: Artdata

Atlantic Reference Centre Museum of Canadian Atlantic Organisms - Invertebrates and Fishes Data

Australian Museum: Australian Museum provider for OZCAM

Buschbaum, Christian (2010): Abundance of macrobenthos organisms in the northern Wadden Sea in 2007. Alfred Wegener Institute for Polar and Marine Research - Wadden Sea Station Sylt, doi:10.1594/PANGAEA.755036

Buschbaum, Christian (2010): Abundance of macrobenthos organisms in the northern Wadden Sea in 2010. Alfred Wegener Institute for Polar and Marine Research - Wadden Sea Station Sylt, doi:10.1594/PANGAEA.755039

Buschbaum, Christian (2010): Abundance of macrobenthos organisms in the northern Wadden Sea in 2008. Alfred Wegener Institute for Polar and Marine Research - Wadden Sea Station Sylt, doi:10.1594/PANGAEA.755037

Buschbaum, Christian (2010): Abundance of macrobenthos organisms in the northern Wadden Sea in 2009. Alfred Wegener Institute for Polar and Marine Research - Wadden Sea Station Sylt, doi:10.1594/PANGAEA.755038

California Academy of Sciences: CAS Invertebrate Zoology (IZ)

City of Amsterdam: City of Amsterdam (NL) - Fyke Fishing Piet Ruiter

Craeymeersch, Johan A; Duineveld, Gerard C A (2011): Abundance of benthic infauna in surface sediments from the North Sea sampled during cruise Tridens00/5. doi:10.1594/PANGAEA.756784

Department of Biology, University of Copenhagen: Marine Benthic Fauna List, Island of Læsø, Denmark

Department of Biology, University of Copenhagen: Nivå Bay species list, Zealand, Denmark

Diveboard - Scuba diving citizen science observations. Online at and

Dutch Foundation for Applied Water Research : Dutch Foundation for Applied Water Research (STOWA) - Limnodata Neerlandica

EAISSNA - An Electronic Atlas of Ichthyoplankton on the Scotian Shelf of North America

EMAP requests that all individuals who download EMAP data acknowledge the source of these data in any reports, papers, or presentations. If you use these data, please include a statement similar to "Some or all of the data described in this article were produced by the U.S. Environmental Protection Agency through its Environmental Monitoring and Assessment Program (EMAP),

Field Museum: Field Museum of Natural History (Zoology) Invertebrate Collection

Florida Museum of Natural History: invertebratezoology

Fockedey N.; Beyst, B.; Cattrijsse, A.; Dewicke A.; Deneudt, K.; Mees J.; Vincx, M.(2004). Historical hyperbenthos data (1987-2001) from the North Sea and some adjacent areas. Collaboration between Ghent University (UGent), Biology Department, Marine Biology Section and Flanders Marine Institute (VLIZ). Data consulted through EurOBIS on dd/mm/yyyy.

Frauenheim, K et al. (2009): Benthos counted on dredged samples during Valdivia cruise VA44. doi:10.1594/PANGAEA.717137, In Supplement to: Frauenheim, Karin; Neumann, V; Thiel, Hjalmar; Türkay, Michael (1989): The distribution of the larger epifauna during summer and winter in the North Sea and its suitability for environmental monitoring. Senckenbergiana maritima, 20(3/4), 101-118,

GEO-Tag der Artenvielfalt: Artenvielfalt auf den Elbwiesen (Dessau)

GEO-Tag der Artenvielfalt: Artenvielfalt der Nordsee - Bremerhaven (Dorum-Neufeld)

GEO-Tag der Artenvielfalt: Artenvielfalt der Nordsee - Helgoland

GEO-Tag der Artenvielfalt: Artenvielfalt der Nordsee - Sylt

GEO-Tag der Artenvielfalt: Gesamtartenliste Bremerhaven, Helgoland und Sylt

GEO-Tag der Artenvielfalt: Schatzinsel Norderney

GEO-Tag der Artenvielfalt: Tauchen und Meer 02

GEO-Tag der Artenvielfalt: Wangerooge

GEO-Tag der Artenvielfalt: Wattenmeer-Safari (Wurster Watt)

GEO-Tag der Artenvielfalt: Wattuntersuchung (Cuxhaven)

GEO-Tag der Artenvielfalt: Wulfsmuehle/Pinnau

GEO-Tag der Artenvielfalt: Zukünftiges NSG Höftland/Bockholmwik

ICoMM MICROBIS

iNaturalist.org: iNaturalist research-grade observations

Kochmann, J; Reise, K (2008): Endofauna abundance in 2006-10 from an experiment in the Wadden Sea of List, Sylt. Alfred Wegener Institute for Polar and Marine Research - Wadden Sea Station Sylt, doi:10.1594/PANGAEA.686208, In: Kochmann, Judith; Reise, Karsten (2008): Abundance of organisms during a field experiment in the Wadden Sea off the island of Sylt, Northern Germany. Alfred Wegener Institute for Polar and Marine Research - Wadden Sea Station Sylt, doi:10.1594/PANGAEA.686220

Kochmann, J; Reise, K (2008): Epi-endofauna abundance in 2007-03 from an experiment in the Wadden Sea off List, Sylt. doi:10.1594/PANGAEA.686230, In: Kochmann, Judith; Reise, Karsten (2008): Abundance of organisms during a field experiment in the Wadden Sea off the island of Sylt, Northern Germany. Alfred Wegener Institute for Polar and Marine Research - Wadden Sea Station Sylt, doi:10.1594/PANGAEA.686220

Kochmann, J; Reise, K (2008): Epifauna abundance in 2006-10 from an experiment in the Wadden Sea of List, Sylt. Alfred Wegener Institute for Polar and Marine Research - Wadden Sea Station Sylt, doi:10.1594/PANGAEA.686209, In: Kochmann, Judith; Reise, Karsten (2008): Abundance of organisms during a field experiment in the Wadden Sea off the island of Sylt, Northern Germany. Alfred Wegener Institute for Polar and Marine Research - Wadden Sea Station Sylt, doi:10.1594/PANGAEA.686220

Laborda, A.J. (2007) CRAI-Experimental, Colecciones Zoológicas ULE, Colección de Malacostráceos.

Marine Invertebrate Diversity Initiative (MIDI) Database

Museum Victoria: Museum Victoria provider for OZCAM

National Biodiversity Data Centre: BioMar - LIFE project

National Biodiversity Data Centre: Ireland's BioBlitz 2012

National Biodiversity Data Centre: Marine species distributions in Irish coastal waters

National Biodiversity Data Centre: Sponges of Rathlin Island

National geological collection of Estonia: University of Tartu, Museum of Geology

Natural History Museum, University of Oslo: BioFokus

Naturalis Biodiversity Center: Naturalis National Natural History Museum (NL) – Crustacea_Decapoda

Netherlands Biodiversity Information Facility (NLBIF): Zoological Museum Amsterdam, University of Amsterdam (NL) – Benthos monitoring of the North Sea_ research database

NIOZ Royal Netherlands Institute for Sea Research: Royal Netherlands Institute for Sea Research (NIOZ) - Kom Fyke Mokbaai

Ocean Biogeographic Information System: A comparison of benthic biodiversity in the North Sea, English Channel and Celtic Seas (EurOBIS)

Ocean Biogeographic Information System: Biogeography Scheldt Estuary (EurOBIS)

Ocean Biogeographic Information System: MarBEF Publication Series data (EurOBIS)

Pohle, G., L. Van Guelpen, A. Martin, D. Welshman, and A. McGuire. 2004. Bay of Fundy Species

Prince, Philip (2011): Abundance of benthic infauna in surface sediments from the North Sea sampled during cruise Dana00/5. doi:10.1594/PANGAEA.756781

Queen Victoria Museum and Art Gallery: Queen Victoria Museum Art Gallery provider for OZCAM

Rigby,P.R., B.Konar, T.Kato, K.Iken, H.Chenelot and Y.Shirayama (2005)NaGISA OBIS Dataset ver.1

Royal Belgian Institute of Natural Sciences: RBINS collections

Senckenberg: Collection Crustacea - SNSD

Senckenberg: Collection Crustacea - ZIM Hamburg

Senckenberg: Collection Crustacea - ZMB

Senckenberg: Collection Crustacea SMF

Senckenberg: Crustacea ZMK

Senckenberg: Insecta ZMK

SPN - Service du Patrimoine naturel, Muséum national d'Histoire naturelle, Paris: Inventaire National du Patrimoine Naturel : Données Benthos du Réseau des Stations et Observatoires Marins

SPN - Service du Patrimoine naturel, Muséum national d'Histoire naturelle, Paris: Inventaire National du Patrimoine Naturel : Crustacés Décapodes de France

Stevens, M. et al. (2009). Onderzoek naar de trekvissoorten in het stroomgebied van de Schelde. Rapport van het Instituut voor Natuur- en Bosonderzoek, R.2009.9. Instituut voor Natuur- en Bosonderzoek: Brussel, Belgium. 188 pp.

The Norwegian Biodiversity Information Centre (NBIC): Norwegian Species Observation Service

UK National Biodiversity Network: Bristol Regional Environmental Records Centre - BRERC October 2009

UK National Biodiversity Network: Centre for Environmental Data and Recording - Marine Data from Northern Ireland

UK National Biodiversity Network: Countryside Council for Wales - Pembrokeshire Marine Species Atlas

UK National Biodiversity Network: Countryside Council for Wales - Marine data from Countryside Council for Wales (CCW) Technical Support (Research & Monitoring) Contracts, Wales

UK National Biodiversity Network: Countryside Council for Wales - 1996 - 2005 CCW Marine Intertidal Phase 1 species dataset

UK National Biodiversity Network: Countryside Council for Wales - Welsh Invertebrate Database (WID)

UK National Biodiversity Network: Cumbria Biodiversity Data Centre - Cumbria Biodiversity Data Centre. Invertebrate Observation Records other than Lepidoptera. Pre-2010 for Cumbria

UK National Biodiversity Network: Cumbria Biodiversity Data Centre - Norman and Florence Hammond records. Seawatch and coastal survey records.

UK National Biodiversity Network: Highland Biological Recording Group - HBRG Other Invertebrates Dataset

UK National Biodiversity Network: Isle of Wight Local Records Centre - IOW Natural History & Archaeological Society Marine Invertebrate Records 1853- 2011

UK National Biodiversity Network: Joint Nature Conservation Committee - Marine Nature Conservation Review (MNCR) and associated benthic marine data held and managed by JNCC

UK National Biodiversity Network: Marine Biological Association - Volunteer sightings data held by the DASSH Data Archive Centre

UK National Biodiversity Network: Marine Biological Association - Marine Life Survey Data (collected by volunteers) collated by MarLIN

UK National Biodiversity Network: Marine Biological Association - Marine survey data (Professional) held by <i>MarLIN</i>.

UK National Biodiversity Network: Marine Biological Association - DASSH Data Archive Centre Academic surveys

UK National Biodiversity Network: Marine Biological Association - DASSH Data Archive Centre expert sighting records

UK National Biodiversity Network: Marine Conservation Society - Seasearch Marine Surveys

UK National Biodiversity Network: Merseyside BioBank - Merseyside BioBank Active Naturalists (unverified)

UK National Biodiversity Network: Merseyside BioBank - North Merseyside Other Taxa (unverified)

UK National Biodiversity Network: Natural England - Marine Nature Conservation Review (MNCR) and associated benthic marine data held and managed by English Nature

UK National Biodiversity Network: North & East Yorkshire Ecological Data Centre - North and East Yorkshire Ecological Data Centre - Non-sensitive Records from all taxonomic groups.

UK National Biodiversity Network: Porcupine Marine Natural History Society - Marine flora and fauna records from the North-east Atlantic

UK National Biodiversity Network: Scottish Natural Heritage - Marine Nature Conservation Review (MNCR) and associated benthic marine data held and managed by Scottish Natural Heritage

UK National Biodiversity Network: South East Wales Biodiversity Records Centre - CCW Regional Data : South East Wales Non-sensitive Species Records

UK National Biodiversity Network: Suffolk Biological Records Centre - Suffolk Biological Records Centre (SBRC) dataset

UK National Biodiversity Network: Yorkshire Naturalists' Union Marine and Coastal Section - Yorkshire Naturalists Union Marine and Coastal Section Records

UK National Biodiversity Network: Yorkshire Wildlife Trust - Yorkshire Wildlife Trust Shoresearch

USGS/NAS database. 2012. Gainesville, FL. USGS/NAS database

Vanden Berghe, E. (2000). Tisbe, Taxonomic Information System for the Belgian coastal area.

VIS Fishes in inland waters in Flanders, Belgium dataset (2013); Research Institute for Nature and Forest(INBO). Online: VIS Fishes in inland waters in Flanders, Belgium dataset

GBIF search term “Charybdis hellerii”

Australian Museum: Australian Museum provider for OZCAM

Chavan, VIshwas and C. T. Achuthankutty (editors), IndOBIS Catalogue of Life, Available at Retrived day, date, year

Florida Museum of Natural History: invertebratezoology

GBIF-Spain: Colección de Crustáceos Decápodos y Estomatópodos del Centro Oceanográfico de Cádiz: CCDE-IEOCD

Marine Benthos - BIOTA/FAPESP

MNHN - Museum national d'Histoire naturelle: Crustaceans specimens

Museum Victoria: Museum Victoria provider for OZCAM

National Museum of Natural History, Smithsonian Institution: NMNH occurrence DwC-A

Ocean Biogeographic Information System: The SERTC Invertebrate Database: Invertebrates of the southeastern United States

Senckenberg: Collection Crustacea - ZMB

Senckenberg: Collection Crustacea SMF

GBIF search term “Charybdis japonica”

National Institute of Genetics, ROIS: Marine Invertebrata specimen database of Osaka Museum of Natutal History

National Museum of Natural History, Smithsonian Institution: NMNH occurrence DwC-A

National Museum of Nature and Science, Japan: Akita Prefectural Museum＋Malacostracans Specimens of Akita Prefectural Museum

Ocean Biogeographic Information System: NCOS1959_Crustacea (OBIS China)

Senckenberg: Collection Crustacea - ZMB

GBIF search term “Hemigrapsus sanguineus”

iNaturalist.org: iNaturalist research-grade observations

National Institute of Genetics, ROIS: Marine Invertebrata specimen database of Osaka Museum of Natutal History

Senckenberg: Collection Crustacea - ZMB

Senckenberg: Collection Crustacea SMF

USGS/NAS database. 2012. Gainesville, FL. USGS/NAS database

Vanden Berghe, E. (2000). Tisbe, Taxonomic Information System for the Belgian coastal area.

GBIF search term “Rhithropanopeus harrisii”

Atlantic Reference Centre Museum of Canadian Atlantic Organisms - Invertebrates and Fishes Data

Batllori Sampedro, E., Avilés Ramírez, G. y H. Medina Quijano. 2007. Biodiversidad de la macrofauna bentónica en la microcuenca de Chabihau, Yucatán. Centro de Investigación y de Estudios Avanzados del Instituto Politécnico Nacional. Unidad Mérida. Bases de datos SNIB2010-CONABIO proyecto No. CE021. México, D.F.

California Academy of Sciences: CAS Invertebrate Zoology (IZ)

City of Amsterdam: City of Amsterdam (NL) - Fyke Fishing Piet Ruiter

CNCR, Portal UNIBIO, Instituto de Biología, Universidad Nacional Autónoma de México, consultada el dd/mm/yy.

Diveboard - Scuba diving citizen science observations. Online at and

Dutch Foundation for Applied Water Research : Dutch Foundation for Applied Water Research (STOWA) - Limnodata Neerlandica

GEO-Tag der Artenvielfalt: GEO-Hauptveranstaltung Bodden (Vilm)

Institute of Nature Conservation, Polish Academy of Sciences: Alien Species in Poland - Animals

León García T. J., Herrera Castillo, E. G. y J. L. Hernández Aguilera. 2006. Actualización de la base de datos de crustáceos y registro de datos de peces del pacífico mexicano de la Colección Biológica de la Secretaría de Marina. Dirección General Adjunta de Oceanografía, Hidrografía y Meteorología. Secretaría de Marina. Bases de datos SNIB2010-CONABIO. Crustáceos. Proyectos No. BA007, B035 y P001. México, D.F.

M.E.Conkright, J.I. Antonov, O. Baranova, T.P. Boyer, H.E. Garcia, R. Gelfeld, D. Johnson, R.A. Locarnini, P.P., T.D. O'Brien, I. Smolyar, C. Stephens, 2002: World Pcean Database 2001, Voulme 1: Introduction. Ed: Sydney Levitus, NOAA Atlas NESDIS 42, U.S. Government Printing Office, Washinton, D.C., 167 pp.

Museum of Comparative Zoology, Harvard University: Museum of Comparative Zoology, Harvard University

National Museum of Natural History, Smithsonian Institution: NMNH occurrence DwC-A

Ocean Biogeographic Information System: Biogeography Scheldt Estuary (EurOBIS)

Ocean Biogeographic Information System: MarBEF Publication Series data (EurOBIS)

Ocean Biogeographic Information System: The SERTC Invertebrate Database: Invertebrates of the southeastern United States

Rigby,P.R., B.Konar, T.Kato, K.Iken, H.Chenelot and Y.Shirayama (2005)NaGISA OBIS Dataset ver.1

Senckenberg: Collection Crustacea - ZMB

Senckenberg: Collection Crustacea SMF

U.S. Environmental Protection Agency: Environmental Monitoring and Assessment Program (EMAP),

USGS/NAS database. 2012. Gainesville, FL. USGS/NAS database

Vanden Berghe, E. (2000). Tisbe, Taxonomic Information System for the Belgian coastal area.

Supplemental Literature Review: Citations for occurrence records

Ahl, R. S. & S. P. Moss, 1999. Status of the nonindigenous crab, Hemigrapsus sanguineus, at Greenwich Point, Connecticut. Northeastern Naturalist 6: 221-224.

Akiyama, S., E. Saito & T. Watanabe, 2007. Relationship between soak time and number of enmeshed animals in experimentally lost gill nets. Fisheries Science (Tokyo) 73: 881-888.

Almeida, M. J., J. Ignacio Gonzalez-Gordillo, A. A. V. Flores & H. Queiroga, 2011. Cannibalism, post-settlement growth rate and size refuge in a recruitment-limited population of the shore crab Carcinus maenas. Journal of Experimental Marine Biology and Ecology 410: 72-79.

Archdale, M. V., C. P. Anasco & S. Hiromori, 2006. Comparative fishing trials for invasive swimming crabs Charybdis japonica and Portunus pelagicus using collapsible pots. Fisheries Research (Amsterdam) 83: 50-55.

Bernadochi, L. C., F. Silvestri & A. Turra, 2012. Composition and seasonal variation of Brachyura and Anomura (Crustacea, Decapoda) associated with brown mussel farms at Praia da Cocanha, Brazil. Brazilian Archives of Biology and Technology 55: 549-557.

Blakeslee, A. M. H., C. H. McKenzie, J. A. Darling, J. E. Byers, J. M. Pringle & J. Roman, 2010. A hitchhiker's guide to the Maritimes: anthropogenic transport facilitates long-distance dispersal of an invasive marine crab to Newfoundland. Diversity and Distributions 16: 879-891.

Bolanos, J. A., J. Antonio Baeza, J. E. Hernandez, C. Lira & R. Lopez, 2012. Population dynamics and reproductive output of the non-indigenous crab Charybdis hellerii in the south-eastern Caribbean Sea. Journal of the Marine Biological Association of the United Kingdom 92: 469-474.

Breton, G., M. Faasse, P. Noel & T. Vincent, 2002. A new alien crab in Europe: Hemigrapsus sanguineus (Decapoda: Brachyura: Grapsidae). Journal of Crustacean Biology 22: 184-189.

Burden, C. T., A. J. Stow, S. J. Hoggard, M. A. Coleman & M. J. Bishop, 2014. Genetic structure of Carcinus maenas in southeast Australia. Marine Ecology Progress Series 500: 139-147.