Table 1. Review of hadrosaurid specimens previously referred to Kritosaurus. Geochronological ages after the following sources: Aguja Fm (Sankey 2005; Sullivan & Lucas 2006); Bearpaw Fm (Toraryk 1987); Cerro del Pueblo Fm (Eberth et al. 2004; Kirkland et al. 2006); Dinosaur Park Fm (Eberth 2005); El Picacho Fm (Lehman 1996, 2001); Fruitland Fm (Sullivan & Lucas 2006); Javelina Fm (Lehman et al. 2006; Hunt & Lehman 2008); Judith River Fm (Eberth 1997); Kirtland Fm (Sullivan & Lucas 2006); Los Alamitos Fm (Agnolin & Martinelli 2009; Prieto-Márquez & Salinas 2010); Milk River Fm (Payenberg et al. 2002); Two Medicine Fm (Horner 1992; Trexler 2001); Olmos Fm (Kirkland et al. 2006); and Woodbury Fm (Litwin et al. 1993). Locality information follows the literature sources containing the previous Kritosaurus referrals listed below.
Specimen / Material / Locality / Horizon / Previous Referral / Revised Referral / Justification for Revised Referral /AMNH 3079 / Right maxilla, ulna, scapula, coracoid, and skull fragments / San Vicente, Big Bend National Park, TX, USA / Upper shale member of Aguja Fm (late Campanian-early Maastrichtian) / ?Kritosaurus cf. K. navajovius (Davies 1983; Wagner 2001) / Saurolophinae / Lacking Kritosaurus diagnostic characters /
AMNH 5350 / Skull roof with braincase, partial postcranium / Little Sandhill Creek, Dinosaur Provincial Park, AB, Canada / Dinosaur Park Fm (late Campanian) / Kritosaurus cf. notabilis (Lull & Wright 1942; Wagner 2001) / Gryposaurus notabilis / Horner (1992); Prieto-Márquez (2010b) /
Table 1. (Continued)
Specimen / Material / Locality / Horizon / Previous Referral / Revised Referral / Justification for Revised Referral /AMNH 5465 / Skull elements, partial postranium / SW of Cut Bank, Glacier County, NW MT, USA / Lower Two Medicine Fm (early Campanian) / Kritosaurus (Brett-Surman 1989; Brett-Surman & Wagner 2007); Kritosaurus latidens (Wagner 2001) / Gryposaurus latidens / Horner (1992); Prieto-Márquez (2012) /
AMNH 5797 / Left maxilla, dentary coronoid process, and other fragments / Kimbetoh Wash, San Juan County, NM, USA / Naashoibito Member of Kirtland Fm (Maastrichtian) / Kritosaurus navajovius (Sinclair & Granger 1914; Lucas et al. 1987); ‘Hadrosaurinae’(Williamson 2000) / Saurolophinae / Williamson, (2000); lacking Kritosaurus diagnostic characters /
AMNH 5799 / Partial skull and atlas / Ojo Alamo, San Juan County, NM, USA / De-Na-Zin Member of Kirtland Fm (late Campanian) / Kritosaurus navajovius (Brown 1910) / Kritosaurus navajovius / Brown (1910); holotype (see revised diagnosis, this paper) /
AMNH 8525 / About a dozen teeth / Clam Bed Hollow, Chouteau County, MT, USA / Judith River Fm (middle-late Campanian) / Kritosaurus cf. K. breviceps (Sahni 1972) / Hadrosauridae / Lacking Kritosaurus diagnostic characters /
ANSP 10005 / Maxillary fragment and partial postcrania / Haddonfield, NJ, USA / Woodbury Fm (early Campanian) / Hadrosaurus (Leidy 1858), Kritosaurus (Baird & Horner 1977) / Hadrosaurus / Leidy (1858); Prieto-Márquez et al. (2006); Prieto-Márquez (2011) /
Table 1. (Continued)
Specimen / Material / Locality / Horizon / Previous Referral / Revised Referral / Justification for Revised Referral /BYU 12950 / Partial articulated skull (see main text for details) / NW flank of Betonnie Tsosie Wash, San Juan County, NM, USA / Farmington Member of Kirtland Fm (late Campanian) / Kritosaurus navajovius (Horner 1992; Williamson 2000; Wagner 2001; Prieto-Márquez 2010a), Anasazisaurus horneri (Hunt & Lucas 1993; Lucas et al. 2006) / Kritosaurus horneri / BYU 12950 cannot be distinguished from Kritosaurus navajovius holotype when considering overlapping elements. Non-overlapping elements in BYU 12950 show autapomorphies /
CMN 2278 / Nearly complete skull and partial postcrania / Near Reed Deer River, Dinosaur Provincial Park, AB, Canada / Dinosaur Park Fm (late Campanian) / Gryposaurus notabilis (Lambe 1914), Kritosaurus notabilis (Brett-Surman 1989; Wagner 2001) / Gryposaurus notabilis / Lambe (1914); Horner (1992); Gates & Sampson (2007) /
GSC 8729 / Various tooth crowns / Deadhorse Coulee, near Milk River, adjacent to Sweetgrass hills, AB, Canada / Upper Milk River Fm (late Santonian-earliest Campanian) / cf. Kritosaurus sp. (Russell 1935) / Hadrosauridae / Lacking Kritosaurus diagnostic characters /
Table 1. (Continued)
Specimen / Material / Locality / Horizon / Previous Referral / Revised Referral / Justification for Revised Referral /IGM 6685 / Partial premaxilla, dentary, and predentary / 1 km N of Presa de San Antonio, central Parras Basin, Coahuila, Mexico / Cerro del Pueblo Fm (latest Campanian) / Kritosaurus sp. (Serrano-Brañas 2006; Kirkland et al. 2006) / Kritosaurus navajovius / Possession of diagnostic combination of Kritosaurus navajovius mandibular characters. Specimen cannot be distinguished from K. navajovius when considering overlapping elements /
LSUMG V-1183 / Right quadrate / Dogie Mountain, USGS Tule Mountain 7.5" quadrangle, Texas / Uppermost Javelina Fm (middle-late Maastrichtian) / ?Kritosaurus cf. K. navajovius (Wagner 2001) / Saurolophinae / Lacking Kritosaurus diagnostic characters /
Table 1. (Continued)
Specimen / Material / Locality / Horizon / Previous Referral / Revised Referral / Justification for Revised Referral /MACN-RN 2 / Right prefrontal, left postorbital, partial sacrum, caudal vertebrae, sternals, coracoid, scapula, ilia, ischia, pubes, and femora / Arroyo Verde region, SE of Rio Negro Province, Argentina / Los Alamitos Fm (late Campanian-early Maastrichtian) / Kritosaurus australis (Bonaparte et al. 1984, 1996; Bonaparte & Rougier 1987) / Secernosaurus koerneri / Wagner (2001); Prieto-Márquez & Salinas (2010) /
MACN-RN 142 / Partial braincase with parietal, frontal, and postorbita, partial predentary and dentaries, right maxilla, and left scapula / Arroyo Verde region, SE of Rio Negro Province, Argentina / Los Alamitos Fm (late Campanian-early Maastrichtian) / Kritosaurus australis (Bonaparte et al. 1984, 1996; Bonaparte & Rougier 1987) / Secernosaurus koerneri / Wagner (2001); Prieto-Márquez & Salinas (2010) /
MACN-RN 143 / Partial braincase / Arroyo Verde region, SE of Rio Negro Province, Argentina / Los Alamitos Fm (late Campanian-early Maastrichtian) / Kritosaurus australis (Bonaparte et al. 1984, 1996; Bonaparte & Rougier 1987) / Secernosaurus koerneri / Wagner (2001); Prieto-Márquez & Salinas (2010) /
Table 1. (Continued)
Specimen / Material / Locality / Horizon / Previous Referral / Revised Referral / Justification for Revised Referral /MACN-RN 144 / Partial skull roof and braincase / Arroyo Verde region, SE of Rio Negro Province, Argentina / Los Alamitos Fm (late Campanian-early Maastrichtian) / Kritosaurus australis (Bonaparte et al. 1984, 1996; Bonaparte & Rougier 1987) / Secernosaurus koerneri / Wagner (2001); Prieto-Márquez & Salinas (2010) /
MACN-RN 145 / Left radius and ulna, distal femur, left metatarsals II and III, and middle dorsal vertebrae / Arroyo Verde region, SE of Rio Negro Province, Argentina / Los Alamitos Fm (late Campanian-early Maastrichtian) / Kritosaurus australis (Bonaparte et al. (1984, 1996; Bonaparte & Rougier 1987) / Secernosaurus koerneri / Wagner (2001); Prieto-Márquez & Salinas (2010) /
MACN-RN 146 / Right scapula, partial ilia, and proximal left ischium / Arroyo Verde region, SE of Rio Negro Province, Argentina / Los Alamitos Fm (late Campanian-early Maastrichtian) / Kritosaurus australis (Bonaparte et al. 1984, 1996; Bonaparte & Rougier 1987) / Secernosaurus koerneri / Wagner (2001); Prieto-Márquez Salinas, (2010) /
MNA Pl. 1626 / Maxillary and dentary teeth / near Burnham, San Juan County, NM, USA / Upper Fruitland Fm (late Campanian) / Kritosaurus navajovius (Armstrong-Ziegler 1980; Lucas et al. 1987) / Hadrosauridae / Lacking Kritosaurus diagnostic characters /
MOR 399 / Nasal, lacrimal, pterygoid, quadrate, and humerus / MOR locality JR-027, Goffena Ranch, Delphia, Musselshell County, MT, USA / Judith River Fm (middle-late Campanian) / Gryposaurus incurvimanus (Horner 1992); Kritosaurus notabilis (Wagner 2001) / Gryposaurus cf. notabilis / Prieto-Márquez, (2010b) /
Table 1. (Continued)
Specimen / Material / Locality / Horizon / Previous Referral / Revised Referral / Justification for Revised Referral /MOR 478 / Cranial and postcranial elements from at least two individuals including nasal, maxilla, jugals, squamosal, surangulars, and axial, pectoral, pelvic and limb bones / MOR locality TM-055, Hillside Quarry bonebed at Harwood Cliff Site, Pondera County, NW MT, USA / Lower Two Medicine Fm (early Campanian) / Kritosaurus latidens (Wagner 2001) / Gryposaurus latidens / Horner (1992); Prieto-Márquez (2012) /
MSNM V345 / Nearly complete skull and postcrania / Near Steveville in Red Deer River Valley, Dinosaur Provincial Park, AB, Canada / Dinosaur Park Fm (late Campanian) / Kritosaurus notabilis (Vialli 1960; Pinna 1979; Wagner 2001) / Gryposaurus notabilis / Prieto-Márquez (2010b) /
NMMNH P-16106 / Partial articulated skull with braincase, humerus, and vertebrae (see main text) / NMMNH locality L-346, Willow Wash, San Juan County, NM, USA / De-Na-Zin Member of Kirtland Fm (late Campanian) / Kritosaurus navajovius (Horner 1992; Williamson 2000; Wagner 2001; Prieto-Márquez 2010a), Naashoibitosaurus ostromi (Hunt & Lucas 1993) / Naashoibitosaurus ostromi / Lacking of Kritosaurus diagnostic characters and possession of autapomorphic characters /
Table 1. (Continued)
Specimen / Material / Locality / Horizon / Previous Referral / Revised Referral / Justification for Revised Referral /NMMNH P-25057 / Partial sacrum, right ilium, partial pubes, left femur, left tibia, fibulae, metatarsals, and vertebrae / NMMNH locality 3117, San Juan Basin, NM, USA / Fruitland/basal Kirtland Fm (late Campanian) / Kritosaurus navajovius (Wagner 2001) / Saurolophinae / Lacking Kritosaurus diagnostic characters /
NMMNH P-31928 / Left quadrate, left humerus, and ilium / NMMNH locality 4104, San Juan Basin, NM, USA / Fruitland/basal Kirtland Fm (late Campanian) / Kritosaurus navajovius (Wagner 2001) / Saurolophinae / Williamson, 2000; Lacking Kritosaurus diagnostic characters /
OMNH 8-21-S6 and S10 / Two femora / Unknown to the author / Upper shale member of Aguja Fm (late Campanian-early Maastrichtian) / Kritosaurus cf. K. navajovius (Wagner 2001) / Saurolophinae / Lacking Kritosaurus diagnostic characters /
PASAC-1 / Partial cranial and postcranial elements / near El Mezquite, 4 km SW of Sabinas, Coahuila, Mexico / lower Olmos Fm (latest Campanian) / Kritosaurus sp. (Serrano-Brañas 2006; Kirkland et al. 2006) / Saurolophinae / Lacking Kritosaurus diagnostic characters /
PMU.R93 (not R245, Mateer, 1981) / Left sternal / South flank of Kimbetoh, San Juan County, NM, USA / Kirtland Formation / Kritosaurus cf. K. navajovius (Mateer 1981); ‘Hadrosaurinae’ (Williamson 2000) / Saurolophinae / Lacking Kritosaurus diagnostic characters /
Table 1. (Continued)
Specimen / Material / Locality / Horizon / Previous Referral / Revised Referral / Justification for Revised Referral /PMU.R94 (not R224, Mateer, 1981) / Partial left pubis / South flank of Kimbetoh, San Juan County, NM, USA / Kirtland Formation / Kritosaurus cf. K. navajovius (Mateer 1981); ‘Hadrosaurinae’ (Williamson 2000) / Saurolophinae / Lacking Kritosaurus diagnostic characters /
PMU.R95 (not R236, Mateer, 1981) / Left ischium / South flank of Kimbetoh, San Juan County, NM, USA / Kirtland Formation / Kritosaurus cf. K. navajovius (Mateer 1981); ‘Hadrosaurinae’ (Williamson 2000) / Saurolophinae / Lacking Kritosaurus diagnostic characters /
PMU.R96 (not R223, Mateer, 1981) / Partial right humerus / South flank of Kimbetoh, San Juan County, NM, USA / Kirtland Formation / Kritosaurus cf. K. navajovius (Mateer 1981); ‘Hadrosaurinae’ (Williamson 2000) / Saurolophinae / Lacking Kritosaurus diagnostic characters /
PMU.R216 (not R231, Mateer, 1981) / Right ilium / South flank of Kimbetoh, San Juan County, NM, USA / Kirtland Formation / Kritosaurus cf. K. navajovius (Mateer 1981); ‘Hadrosaurinae’ (Williamson 2000) / Saurolophinae / Lacking Kritosaurus diagnostic characters /
ROM 764 / Partially complete skull and nearly complete postcrania / 100 m above Red Deer Rver, 2.5 miles above Happy Jerry ferry, near Sand Hill Creek, AB, Canada / Dinosaur Park Fm (late Campanian) / Kritosaurus incurvimanus (Parks 1919; 1920); Gryposaurus incurvimanus (Horner 1992; Gates & Sampson 2007); Kritosaurus notabilis (Wagner 2001) / Gryposaurus notabilis / Prieto-Márquez (2010b) /
Table 1. (Continued)
Specimen / Material / Locality / Horizon / Previous Referral / Revised Referral / Justification for Revised Referral /ROM 873 / Partially complete articulated skull / Near Steveville in Red Deer River Valley, Alberta, Canada / Dinosaur Park Fm (late Campanian) / Kritosaurus notabilis (Ostrom 1961; Davies 1983; Wagner 2001 as ROM 5859) / Gryposaurus notabilis / Horner (1992); Prieto-Márquez, (2010b) /
TMM 40810 / Ilium / Big Bend National Park, TX, USA / El Picacho Fm (Maastrichtian) / Kritosaurus cf. K. navajovius (Wagner 2001) / Saurolophinae / Lacking Kritosaurus diagnostic characters /
TMM 41442-1 / Left and right maxillae, neural arch and rib fragments / Cow Heaven Mountain, Big Bend National Park, TX, USA / Javelina Fm (middle-late Maastrichtian) / Kritosaurus cf. K. navajovius (Wagner 2001) / Saurolophinae / Lacking Kritosaurus diagnostic characters /
TMM 41917-1 / Left maxilla and teeth / Bruta Canyon, Big Bend National Park, TX, USA / Upper shale member of Aguja Fm (late Campanian-early Maastrichtian) / Kritosaurus cf. K. navajovius (Wagner 2001) / Saurolophinae / Lacking Kritosaurus diagnostic characters /
TMM 41918-3 / Prefrontal and ossified tendons / Bruta Canyon, Big Bend National Park, TX, USA / Upper shale member of Aguja Fm (late Campanian-early Maastrichtian) / Kritosaurus cf. K. navajovius (Wagner 2001) / Saurolophinae / Lacking Kritosaurus diagnostic characters /
TMM 42309-1−11 and 13−18 / Ilia, ischia, pubes, and, ribs, dorsal, sacral, and caudal vertebrae / East of Panther Junction, Big Bend National Park, TX, USA / Upper shale member of Aguja Fm (late Campanian-early Maastrichtian) / ?Kritosaurus (Davies 1983); Kritosaurus cf. K. navajovius (Wagner 2001) / Saurolophinae / Lacking Kritosaurus diagnostic characters /
Table 1. (Continued)
Specimen / Material / Locality / Horizon / Previous Referral / Revised Referral / Justification for Revised Referral /TMM 42325-1 / Left ischium / 19.6 km SW of Glenn Spring, Big Bend National Park, TX, USA / Javelina Fm (middle-late Maastrichtian) / Kritosaurus cf. K. navajovius (Wagner 2001) / Saurolophinae / Davies (1983) /
TMM 42339-3 / Partial left maxilla / Sierra Aguja, Big Bend National Park, TX, USA / upper shale member of Aguja Fm (late Campanian-early Maastrichtian) / ?Kritosaurus cf. K. navajovius (Wagner 2001) / Saurolophinae / Lacking Kritosaurus diagnostic characters /
TMM 42452-1 / Partial skeleton / Rattlesnake Mountain, Big Bend National Park, TX, USA / lower shale member of Aguja Fm / Kritosaurus sp. nov. (Wagner 2001) / Saurolophinae / Study in progress by Wagner /
TMM 42680 / Juvenile right maxilla / Grapevine Hills, Big Bend National Park, TX, USA / upper shale member of Aguja Fm (late Campanian-early Maastrichtian) / Kritosaurus cf. K. navajovius (Wagner 2001) / Saurolophinae / Lacking Kritosaurus diagnostic characters /
TMM 42876 (= 42335-1) / Two dentary teeth / Tule Mountain, northern Dawson Creek, W of Big Bend National Park, TX, USA / Uppermost shale member of Aguja Fm (late Campanian-early Maastrichtian) / Kritosaurus cf. K. navajovius (Wagner 2001) / aff. Kritosaurus navajovius / Tooth crown morphology and denticulation as in K. navajovius /
Table 1. (Continued)