Supplementary material. Studies on the influence of land-use/landscape attributes on plant invasions based on a search in the ISI Web of Science accessed in January 2010. We used the following terms in the search in various combinations: (invas* OR alien) AND (land* structure OR fragmentation OR connectivity OR land-use OR land-cover OR land-use change).

Country/Region / Alien species / Ecosystem / Land-use driver / Landscape attribute / Effect on invasion / Reference
Spain/Macaronesian / several / several / urbanization / dist. to urban areas / ↓ richness / Arévalo et al. 2005
USA/temperate / Lonicera maackii / forest / agricultural intensification / length of corredors / ↑ density / Bartuszevige et al. 2006
dist. to nearest town / ↓ density
USA/temperate / Lonicera maackii / riparian forest / urbanization / % urban land/ / ↑ cover / Borgmann & Rodewald 2005
trail density
% agricultural land / ↓ cover
USA/steppe / Bromus tectorum / shrubland / fragmentation / dist. to roads/powerlines/ / ↓ occurrence / Bradley & Mustard 2006
increased chanelization
USA/temperate / Solanum nigrum, / forest / agricultural intensification / dist. to agricultural land / ↓ cover, richness / Brothers & Spingarm 1992
Chenopodium album,
Taraxacum officinale,
Rosa multiflora
USA/temperate / several / forest / fragmentation / dist. to croplands / = richness / Brothers 1993
USA/temperate / several / forest / fragmentation / dist. to old-fields / ↓ presence / Burke & Nol 1998
Australia/Mediterranean / several / grassland / fragmentation / dist. to urbanized/agricultural land / ↓ cover / Cilliers et al. 2008
South Africa/Mediterranean
USA/temperate / several / grassland / habitat loss / fragment size / = cover, richness / Cully et al. 2003
USA/temperate / several / forest / fragmentation / patch connection / = abundance, richness / Damschen et al. 2006
Belgium/temperate / Prunus serotina / hedgerows / fragmentation / hedgerow intersection / ↑ occurrence / Deckers et al. 2005
Catalonia/Mediterranean / Cortaderia selloana / grassland / urbanization / dist. to urbanized land / ↑ occurrence / Domènech et al. 2005
USA/temperate / several / forested wetland / urbanization / dist. to industrial/commercial land / ↑ richness / Ehrenfeld 2008
USA/semiarid / annuals / wetland / urbanization / lowering water tables / ↑ cover / Elmore et al 2003
USA/temperate / Lonicera maackii, / deciduous forest / transport corridors / dist. to roads / ↓ density / Flory & Clay 2006
Euonymus alata,
Berberis thunbergii,
Pyrus calleryana
USA/temperate / several / forest / fragmentation / dist. to edge / ↓ cover / Fraver 1994
USA/temperate / several / forest / fragmentation / dist. to agricultural/roads/old-fields edges / ↓ cover, richness / Gelhausen et al. 2000
USA/semiarid / several forbs / grassland, shrubland / road improvement / edges in improved roads / ↑ cover, richness / Gelbard & Belnap 2003
Spain/Mediterranean / several / forest / habitat loss / fragment size / ↓ richness / Guirado et al. 2006
dist. to agricultural edge / ↑ richness
dist. to urban edge / = richness
Canada/temperate / perennials / grassland / transport corridors / dist. to roads / = occurrence / Hansen & Clevenger 2005
forest
USA/temperate / Bromus tectorum / several / agricultural intensification / dist. to agricultural land / ↑ cover, richness / Hanson et al. 2008
USA/Mediterranean / several / grassland / habitat loss / patchiness / ↓ richness / Harrison 1999
Belgium/temperate / several / deciduous forest / agricultural intensification / dist. to agricultural land / ↓ cover / Honnay et al. 2002
USA/temperate / Lonicera maackii / deciduous forest / agricultural intensification / connectivity to forested land / ↑ cover / Hutchinson & Vankat 1998
India/tropical / Coffea arabica, / forest / agricultural intensification / dist. to agricultural land / ↓ density / Joshi et al. 2009
C. canephora
USA/steppe / Bromus tectorum / shrubland / habitat loss / transformation to grasslands / ↑ occurrence / Knick & Rotenberry 1997
USA/temperate / Chenopodium album, / grassland, shrubland / infrastructure expansion / disturbed patches / ↑ occurrence / Larson et al. 2001
Agropyron cristatum,
Bromus inermis,
Convolvulus arvensis
Mexico/semiarid / Bromus madritensis / succulent scrub / urbanization / fragments within urban land / ↑ cover / Leyva et al. 2006
Australia/temperate / Rubus fruticosus / forest / aforestation / fragments isolation / ↑ occurrence / Lindenmayer & McCarthy 2001
Pinus radiata
USA/temperate / Lonicera maackii / forest / fragmentation / dist. to anthropic edge / ↓ seedling density / Luken & Goessling 1995
Canada/temperate / several / forest / fragmentation / dist. to edge / ↓ abundance / MacQuarrie & Lacroix 2003
Canada/temperate / Phragmites australis / wetland / infrastructure expansion / managed habitats / ↑ spread / Maheu-Giroux & de Blois 2007
Great Britain/oceanic / several / riparian forest / infrastructure expansion / increased chanelization / ↑ abundance / Maskell et al. 2006
USA/temperate / several / forest / fragmentation / dist. to edge / = diversity / Matlack 1993
Australia/temperate / several / several / fragmentation / road density / ↑ richness / McIntyre & Lavorel 1994
USA/temperate / Lonicera maackii / forest / land abandonment / dist. to edge and streams / ↓ density / Medley 1997
New Zealand/oceanic / several / forest / agricultural intensification / dist. to agricultural land / ↓ richness / Ohlemüller et al. 2006
patch lobation / = richness
fragment size / ↓ richness
pasture area / ↓ richness
forested area / ↓ richness
USA/temperate / Hypericum perforatum / forest / transport networks / road use intensity / ↑ occurrence / Parendes & Jones 2000
Chrysanthemum leucanthemum
Trifolium repens
Hypochaeris radicata
Cirsium vulgare
Chile/subalpine / several / forest / transport networks / dist. to road edge / = richness / Pauchard & Alaback 2004
% pasture in the matrix / ↑ richness
USA/subalpine / several / conifer forest / fragmentation / distance to highways / ↓ cover, richness
USA/temperate / Lonicera spp, / riparian forest / fragmentation / dist. to roads / ↓ occurrence / Predick & Turner 2008
Rhamnus spp
Australia/temperate / several / forest / fragmentation / fragment size / = richness / Ross et al. 2002
New Zealand/temperate / several / ruderal / transport networks / dist. to residential structures/houses / ↓ richness / Sullivan et al. 2009
Germany/temperate / Heracleum mantegazzianum / grassland / fragmentation / habitat connectivity/ / ↑ occurrence / Thiele et al. 2008
habitat shape complexity
USA/temperate / Dioscorea oppositifolia / riparian forest / fragmentation / stream connectivity / ↑ occurrence / Thomas et al. 2006
USA/temperate / several / riparian forest / urbanization / adjacency to commercial malls / ↓ cover, richness / Vidra & Shear 2008
Catalonia/Mediterranean / Opuntia maxima / shrubland / urbanization / dist. to urbanized land / ↓ occurrence / Vilà et al. 2003
O. stricta / woodland / urbanization / dist. to urbanized land / ↓ occurrence
USA/temperate / several / forest / urbanization / dist. to roads / ↓ abundance / Watkins et al. 2003
USA/temperate / Cytisus scoparius, / forest / infrastructureexpansion / stream connectivity to road ditches / ↑ occurrence / Watterson & Jones 2006
Digitalis purpurea
USA/temperate / Rosa multiflorum / forest / agriculture intensification / dist. to agriculture land / ↓ occurrence, density / Weber & Gibson 2007
Ligustrum vulgare / = occurrence, density
New Zealand/oceanic / several / rock outcrop / fragmentation / % grassland in the matrix / ↑ richness / Wiser & Buxton 2008

Supplementary material references not listed in the main text:

Arévalo JR, Delgado JD, Rüdiger Oet al(2005) Distribution of alien vs. native plant species roadside communities along an altitudinal gradient in Tenerife and Gran Canaria (Canary Islands). Persp Plant Ecol Evol Syst7:185-202.

Bradley BA, MustardJF (2006) Characterizing the landscape dynamics of an invasive plant and risk of invasion using remote sensing. Ecol Appl16:1132-1147.

Brothers TS (1993) Fragmentation and edge effects in central Indiana old-growth forests. NatAreas J13:268-275.

Brothers TS, Spingarn A (1992) Forest fragmentation and alien plant invasion of Central Indianaold-growth forests. Conserv Biol6:91-100.

Burke DM, Nol E (1998) Edge and fragment size effects on the vegetation of deciduous forests in Ontario, Canada. NatAreas J 18:45-53.

Elmore AJ, Mustard JF, Manning SJ (2003) Regional patterns of plant community response to changes in water: Owens Valley, California. Ecol Applic13:443-460.

Flory SL, Clay K (2005) Invasive shrub distribution varies with distance to roads and stand age in eastern deciduous forests in Indiana, USA. Plant Ecol184:131-141.

Fraver S(1994) Vegetation responses along edge-to-interior gradients in the mixed hardwood forests of the RoanokeRiver Basin, North Carolina. Conserv Biol 8:822-832.

Gelbard JL, Belnap J (2003) Roads as conduits for exotic plant invasions in a semiarid landscape. Conserv Biol17: 420-432.

Hanson T, Sanchez-de Leon Y, Johnson-Maynard Jet al (2008) Influence of soil and site characteristics on Palouse prairie plant communities. Western North Am Nat68:231-240.

Honnay O, Verheyen K, Hermy M(2002) Permeability of ancient forest edges for weedy plant species invasion. Forest EcolManag161:109-122.

Knick ST, Rotenberry JT (1997)Landscape characteristics of disturbed shrubsteppe habitats in Southwestern Idaho (U.S.A.). Landscape Ecol12:287–297.

Gehlhausen SM,Schwartz NW, Augspurger CK (2000)Vegetation and microclimatic edge effects in two mixed-mesophytic forest fragments. Plant Ecol147:21–35.

Larson DL, Anderson PJ, Newton W(2001) Alien plant invasion in mixed-grass prairie: effects of vegetation type and anthropogenic disturbance. Ecol Appl11:128-141.

Luken JO, Goessling N (1995) Seedling distribution and potential persistence of the exotic shrub Lonicera maackii in fragmented forests. Am Midl Nat 133:124-130.

MacQuarrie K, Lacroix C(2003) The upland hardwood component of prince Edwards Island’s remnant Acadian forest: determination of depth of edge and patterns of exotic plant invasion. Can J Bot81:1113-1128.

Martin PH, Canham CD, Marks PL (2009) Why forests appear resistant to exotic plant invasions: intentional introductions, stand dynamics, and the role of shade tolerance. Front Ecol Environ7:142-179.

Maskell LC, Firbank LG, Thompson K(2006) Interactions between non-native plant species and the floristic composition of common habitats. J Ecol94:1052-1060.

McIntyre S, Lavorel S(1994) Predicting richness of native, rare, and exotic plants in response to habitat and disturbance variables across a variegated landscape. ConservBiol8:521-531.

Matlack GR (1993) Microenvironment variation within and among forest edge sites in the eastern United States. Biol Conserv66:185-194.

Medley KE (1997) Distribution of the non-native shrub Lonicera maackii in Kramer Woods, Ohio. Phys Geogr18:18-36.

Pauchard A, Alaback PB (2006) Edge type defines alien plant species invasions along Pinus contorta burned, highway and clearcut forest edges. Forest Ecol Manag223:327-335.

Predick KI, Turner MG(2008)Landscape configuration and flood frequency influenceinvasive shrubs in floodplain forests of the WisconsinRiver (USA). J Ecol96:91–102.

Ross KA, Fox BJ, Fox MD (2002) Changes to plant species richness in forest fragments: fragment age, disturbance and fire history may be as important as area. J Biogeogr29:749-765.

Thomas JR, Middleton B, Gibson DJ(2006) A landscape perspective of the stream corridor invasion and habitat characteristics of an exotic (Dioscorea oppositifolia) in a pristine watershed in Illinois. Biol Invasions 8:1103-1113.

Watterson NA, Jones JA (2006) Flood and debris flow interactions with roads promote the invasion of exotic plants along steep mountain streams, western Oregon. Geomorphology78:107-123.

Weber JS, Gibson KD (2007) Exotic plant species in old-growth forest in Indiana. Weed Sci55:299-304.

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