Type / Species / References / Formation and age / Location
Arch. / ProtasterpiltonensisSpencer / Spencer (1934a, p. 471, pl. XXXIII, fig. 1) / Upper Mississippian Tournaisian (early Carboniferous) / Pilton, N. Devon, England
Arch. / Protastergranifer (Bohemuragranifer) (Whidborne) / Spencer (1934a, p. 479, pl. XXX, figs 5, 6) / Upper Mississippian Tournaisian (early Carboniferous) / Pilton, N. Devon, England
Arch. / Drepanasterscabrosus(Whidborne) / Spencer (1940, p. 498, pl. XXIX, fig. 6; pl. XXXII, figs 6, 7) / Upper Mississippian Tournaisian (early Carboniferous) / Braunton Down, N. Devon, England
Arch. / Furcasterleptosoma(Salter) / Spencer (1925, p. 319, pl. XVIII, fig. 4; pl. XXII, figs 3–10) / Upper Mississippian
Tournaisian (early Carboniferous) / Braunton Down, N. Devon, England
Arch. / Onychasterbarrisi(Hall) / Spencer (1927, p. 340, pl. XXI, fig. 13) / Upper Mississippian Tournaisian (early Carboniferous) / Braunton Down, N. Devon, England
Arch. / EncrinasterhamlingiSpencer / Spencer (1930, p. 425, pl. XXVII, fig. 10) / Upper Mississippian Tournaisian (early Carboniferous) / Croyde and Braunton, N. Devon, England
Mod. / Aganastergregarius
(Meek & Worthen) / Spencer (1925, p. 285, pl. 20, figs 4, 5);
Jell (1997, p. 312, figs 6.1–6.5) / Border Group, Mississippian Visean (early Carboniferous) / Crawfordsville,
Indiana, USA
Arch. / OnychasterflexilisMeek & Worthen / Spencer (1927, p. 339, pl. XXI, fig. 12) / Upper Mississippian Visean (early Carboniferous) / Skateraw Quarry, Dunbar, Scotland
Arch. / Bohemura(Protaster) granifer / Spencer (1934b, p. 189, pl. VI) / Upper Mississippian
Calcaire du Haut
Viséan(early Carboniferous) / Le HautBanc, near
Boulogne-sur-Mer
France
Mod. / Cholasterpeculiaris(Worthen& Miller) / Spencer & Wright (1966, p. 95, fig. 83.1) / Upper Mississippian (early Carboniferous) / Illinois, USA
Arch. / Stratasterohioensis (KeslingVasseur) / KeslingVasseur
(1971, p. 324, fig. 8) / Early Mississippian (early Carboniferous) / Ohio, USA
Arch. / Vandelooasterplicatilis Jell / Jell (1997, p. 308, pl. 1) / Mississippian (early Carboniferous) / Crawfordsville, Indiana, USA
Arch. / Schoenasterfimbriatus (Meek & Worthen) / Jell (1997, p. 310, pl. 3) / Mississippian (early Carboniferous) / Crawfordsville, Indiana, USA
Mod. / Aganasterjagiellonicus sp. nov. / Thuy et al. (2013, p. 924, fig. 2) / Mississippian - Tournaisian (early Carboniferous) / Southern Poland
Arch. / Protaster sp. / Wilson (1960)
Owen (1965) / Millstone Grit Series Namurian, (early-middle Carboniferous) / Yorkshire, England, UK
Mod. / Syntomospinakuehni Morris et al. / Morris et al. (1973,
p. 474, pl. 1, figs 1–4) / Brush Creek Shale,
Pennsylvanian (late Carboniferous) / Pennsylvania, USA
Arch. / Furcaster sp.[disarticulated vertebrae] / Weller (1930, p.9, pl. 1)
Haude(1982, p.13, pl. 3) / Unnamed Shale below PiasaLimestone, Pennsylvanian(late Carboniferous) / Jersey County, Illinois, USA
Arch. / Furcaster sp. [disarticulated vertebrae] / Weller (1930, p.9, pl. 1)
Lane (1981)
Haude (1982) / Perth Limestone Member
Staunton Formation
Desmoinesian Pennsylvanian(late Carboniferous) / Redwood Creek
Warren County, Indiana, USA
Arch. / Protasteridae sp. [forming
trace fossil Asteriacites
lumbricalis] / Ward (1982)
West & Ward (1990) / Tecumseh Shale Formation, Shawnee Group, Virgillian Pennsylvanian(late Carboniferous) / Wakarusa River,
South of Topeka,
Shawnee County,
Kansas, USA
Arch. / Armathyrasterparadoxis
ophiuroid sp.nov. / Harper & Morris (1978, p. 159, pl. 1) / Brush Creek Shale
Glenshaw Formation
Conemaugh Group Pennsylvanian (late Carboniferous) / Punxsutawney,
Jefferson County, Pennsylvania, USA
Arch. / Furcaster sp.
[as Calclyra sp.] / FrizzellExlein(1955) / Pennsylvanian(late Carboniferous) / Texas, USA
Arch. / Acanthospondyluspennsylvanicus(Harper) / Harper (2014, p. 248) / Glenshaw Formation, Conemaugh Group,Pennsylvanian - early Kasimovian (late Carboniferous) / Pennsylvania, USA
Mod. / Archaeophiomusiumburrisi
(Miller) / Miller (1958, p. 360, fig. 4A);
Miller (1963, p. 725) / Americus Limestone,
early Permian / Augusta, Kansas, USA
Arch. / ?Furcaster[disarticulated arm section] / Kornicker (1954, pl. 6, figs 1a–c, 2a,b)
Imbrie (1955) / Council Grove Group,
Florena Shale Member,
Artinskian(early Permian) / Kansas, USA
Mod. / Archaeophiomusiumburrisi
Zeugophiurinagen. et sp.nov. / Mayou (1967, p. 120, pl. 4) / Loray Formation, Kungurian (early Permian) / Robinson Summit, Nevada, USA
Arch. / Furcasteridaeindet / Perry et al.(2007) / Hueco Group, early Permian / Robledo Mountains, New Mexico, USA
Arch. / Teleosastercreasyi / This paper / Cundlego Formation, early Permian / Southern Carnarvon Basin, Western Australia
Mod. / Archaeophiomusiumburrisi (Miller) / Hattin (1959, p. 1125,
text-fig. 1) / Speiser Shale, early Permian / Southern Kansas, USA
Mod. / Archaeophiomusiumburrisi (Miller) / Hattin (1959, p. 1125,
text-fig. 1);
Hattin (1967, pp. 489–492, text-figs 1–3) / Cottonwood Limestone, early Permian / Oklahoma, USA
Mod. / Archaeophiomusiumburrisi (Miller) / Mayou (1969, p. 936,
pl. 113, fig. 5) / Pequop Formation, early Permian / Ely, Nevada, USA
Mod. / Archaeophiomusium
bispinosumMayou / Mayou (1969, p. 937, pl. 113, figs 1, 2, 4, text-figs 3–5) / PequopFormation, early Permian / Ely, Nevada, USA
Mod. / Archaeophiomusiumandinum
Sanchez / Sanchez (1983, p. 104,
pl. 1, figs 1–3) / PalmaritoFormation, early Permian / Western Venezuela
Arch. / Furcastereiseliana
[as Prosynaptaeiseliana oras Calclyraeiseliana
spine microfossil only] / Spandel(1898, pp. 37–40, pl. XIII); FrizzellExlein(1955); Mostler(1971, p. 8, pl. 11) / Tatarian, Djulfian, Zechsteins,
middle–late Permian / Ranis, Thuringia, Germany
Arch / Furcastergracilis
[as Calclyragracilisspine microfossil only] / KozurMostler(1989, p. 679, pl. 1) / Wordianammonoid-bearingLimestone, Guadalupian(late Permian) / Rupe del Passo de Burgio,Sosio Valley, Sicily, Italy
Arch? / 13 Ophiuroidsindet. / Dunstan (1901) / Lower Bowen, Lower Marine Series, Permian / Leichardt District, central Queensland, Australia
Mod. / Aganaster?fujianensis Liao
& Wang / Liao & Wang (2002,
p. 397, pl. 1, figs 1–10;
pl. 2, figs 1–8) / Tongziyan Formation, Guadalupian
(late Permian) / Fujian, south China
Mod. / Aganaster?huaanensis (Wu) / Wu (1980, p. 61, pl. 1,
figs 1, 2) / TongtzyanFormation, Guadalupian (late Permian) / Fujian, south China
Mod. / Syntomospina?kaiyangensis Feng / Feng (1985, p. 339, pl. 3, figs 3, 4) / LungtanFormation, Wuchiapingian (late Permian) / Guizhou, South China
? / Indeterminate ophiuroid remains / Posenato (1999,
pl. 5, fig. 10) / Tesero Horizon,
Changhsingian(late Permian) / Dolomites, north Italy
? / Disarticulated ossicles / Twitchett Oji (2005) / Griesbachian to Spathian (Early Traissic) / Dolomites, north Italy
Mod. / Huangzhishaniapermotriassica
Chen et al. / Chen et al. (2004, p. 1308, pl. 1, figs 1–6) / ChanghsingFormation, Griesbachian (Early Triassic) / Zhejiang, south China
Mod. / Aplocoma cf. torrii (Desio) / Kummel & Teichert (1970, p. 52, fig. 15) / Kathwai Member,Griesbachian
(Early Triassic) / Salt Range, Pakistan
? / Undescribed body fossils / Twitchettet al. (2005) / Thaynes Formation,
Spathian(Early Triassic) / Nevada, USA
Mod. / Huangzhishaniagulinensis (Feng) / Feng & Gan (1978,
p. 454, pl. 146, fig. 1) / FeixianguanFormation, Olenekian (Early Triassic) / Guizhou, south China
Mod. / Huangzhishaniagulinensis (Feng) / Feng (1985, p. 340, pl. 2, figs 1, 2) / FeixianguanFormation, Olenekian (Early Triassic) / Sichuan, south China
Mod. / Ophiaulaxoidesbijieensis (Feng) / Feng (1985, p. 338, pl. 1, fig. 3) / FeixianguanFormation, Olenekian (Early Triassic) / Guizhou, south China
Mod. / Ophiolepis?shanxiensisYang / Yang et al. (1979, p. 471, pl. 2, figs 17–20) / ShihchienfengFormation, Olenekian (Early Triassic) / Shanxi, north China
Mod. / ‘Ophioderma’? schistovertebrataYang / Yang (1960, p. 159, pl. 1, figs 1, 2) / Feixianguan Formation, Olenekian (Early Triassic) / Guizhou, south China
Mod. / ‘O.’? schistovertebrata Yang / Feng & Gan (1978,
p. 454, pl. 146, fig. 2) / FeixianguanFormation, Olenekian (Early Triassic) / Guizhou, south China
Mod. / PraeaplocomahessiLorigaCavicchi / LorigaCavicchi (1969, pp. 194–196, pls 1–6) / Upper Werfen Formation, Olenekian (Early Triassic) / Dolomites,
north Italy
Mod. / PraeaplocomahessiLorigaCavicchi / Twitchettet al. (2005, p. 214, fig. 2) / Lower Siusi Member, late Griesbachian to Olenekian (Early Triassic) / Dolomites,
north Italy
Mod. / OphiolepisraincsakiDetre / Detre (1983, p. 359, fig. 4) / Campillian Limestone, Olenekian (Early Triassic) / Bakony Mountains, Hungary
Mod. / Aplocoma sp. / Runnegar (1969, pl. 103, figs 16, 17) / Traveston Formation, Smithian (Early Triassic) / Queensland, Australia
Mod. / OphiolepisbalatonicaDetre
Mihaly / DetreMihaly (1987,
p. 449, fig. 1) / Campillian Limestone, Spathian (Early Triassic) / Balaton Highland,
Hungary
Mod. / Ophioderma? haucheconi Eck / Hess (1965, p. 159) / Spathian (Early Triassic) / Berlin, Germany
? / Undescribed body fossils / Twitchettet al. (2005) / Hiraiso Formation,
Smithian(Early Triassic) / Kitakami, Japan
? / Undescribed body fossils / Twitchettet al. (2005) / Elikah Formation,
Smithian(Early Triassic) / Northern Iran
Arch. / ?Eospondylidindet. / Thuy et al. (2017) / Muschelkalk Basin,Anisian(Middle Triassic) / Germany and France
Arch. / ?Eospondylidindet. / Thuy et al. (2017) / Western paleo-Tethys, Anisian (Middle Triassic) / Hungary
Arch. / ?Eospondylidindet. / Thuy et al. (2017, fig. 2) / Eastern paleo-Tethys, Carnian(Middle Triassic) / Yangtse Platform Sichuan, China
Arch. / ?Eospondylidindet. / Thuy et al. (2017) / Western paleo-Tethys,Carnian(Middle Triassic) / Dolomites, north Italy
Arch. / ?Eospondylidindet. / Thuy et al. (2017) / Western paleo-Tethys, Norian(Late Triassic) / Austria
Arch. / ?Eospondylidindet. / Thuy et al. (2017, fig. 2) / paleo-Tethys,Rhaetian(Late Triassic) / Austria
Arch. / ?Eospondylidindet. / Thuy et al. (2017) / Paleo-Tethys, Rhaetian(Late Triassic) / Central Iran
Arch. / ?Eospondylidindet. / Thuy et al. (2017) / South-eastern Tethys, Rhaetian(Late Triassic) / Australia

Supplemental Table 1. Table of all known published Carboniferous, Permian and Triassic ophiuroids (articulated and disarticulated). Data from Chen & McNamara (2006) with further data from. Chen et al. (2004); Detre (1983); DetreMihaly (1987); Dunstan (1901); Feng (1985); Feng & Gan (1978); FrizzellExlein (1955); Harper (2014); Harper & Morris (1978); Hattin (1959); Hattin (1967); Haude (1982); Hess (1965); Imbrie (1955); Jell (1997); KeslingVasseur (1971); Kornicker (1954); KozurMostler (1989); Kummel & Teichert (1970); Lane (1981); Liao & Wang (2002); LorigaCavicchi (1969); Mayou (1967); Mayou (1969); Miller (1958); Miller (1963); Morris et al. (1973); Mostler (1971); Owen (1965); Perry et al. (2007); Posenato (1999); Runnegar (1969); Sanchez (1983); Spandel (1898); Spencer (1925, 1927, 1930, 1934a, b, 1940); Spencer & Wright (1966); Thuy et al. (2015); Thuy et al. (2017); Twitchettet al. (2005); Twitchett & Oji (2005); Ward (1982); Weller (1930); West & Ward (1990); Wilson (1960); Wu (1980); Yang (1960); Yang et al. (1979). Arch. = ‘Archaic’. Mod. = ‘Modern’.

References

Chen, Z. Q., Shi, G. R. Kaiho, K. 2004. New ophiuroids from the Permian/Triassic boundary beds of South China. Palaeontology, 47, 1301–1312.

Detre, C. 1983. Az elsö Ophiuroidea maradvany a magyarországi alsótriászbol. Földtani Közlöny. Bulletin of the Hungary Geological Society, 113, 357–363.

Detre, C. Mihaly, S. 1987. Két Ujabb Ophiuroida Lelet A Balaton-Felvidék Triászából. Magyar Allami Földtani Intézet É vi Jelentese AZ, 1985, 449–452.

Dunstan, B. 1901. [Report on] Geology of the Dawson and Mackenzie Rivers with special reference to the occurrence of Anthracitic coal. Geological Survey of Queensland Reports, 155, 1–28.

Feng, R. L. 1985. New discovery of fossil ophiuroids from Guizhou and southern Sichuan, China. Acta Palaeontologica Sinica, 24, 337–343.

Feng, R. L. Gan, X. M. 1978. Ophiuroidea. Pp. 453–454 in Guizhou Geological and Mineral Resources Bureau. Palaeontological atlas of southwestern China, Guizhou Province. Volume 2. Geological Publishing House, Beijing.

Hattin, D. E. 1959. An occurrence of Ophiuraster burrisi Miller [Kansas]. Journal of Paleontology, 33, 1125–1126.

Hattin, D. E. 1967. Permian ophiuroids from northern Oklahoma. Journal of Paleontology, 41, 489–492.

Haude, R. 1982. Ophiuren (Echinodermata) aus dem Karbon des Rheinischen Schiefergebirges. Geologisches Jahrbuch Hessen, 110, 5–26.

Hess, H. 1965. Trias-Ophiuren aus Deutschland, England, Italien und Spanien. Mitteilungen aus der Bayerischen Staatssammlung für Paläontologie und historische Geologie, 5, 151–177.

Imbrie, J. 1955. Quantitative lithofacies and biofacies study of Florena Shale (Permian) of Kansas. Bulletin of the American Association of Petroleum Geologists, 39, 649–670.

Kornicker, L. S. 1954. Distribution of ostracodes in the Florena Shale. P. 85. Unpublished Master’s essay, Columbia University.

Kozur, H. Mostler, H. 1989. Echinoderm Remains from the Middle Permian (Wordian) from Sosio Valley (Western Sicily). Jahrbuch der Geologischen Bundesanstalt, 132, 677–685.

Kummel, B. Teichert, C. 1970. Stratigraphy and paleontology of the Permian–Triassic boundary beds, Salt Range and trans-Indus ranges, West Pakistan. Pp. 1–110 in B. Kummel & C. Teichert (eds) Stratigraphic boundary problems: Permian and Triassic of West Pakistan, University of Kansas, Geological Department, Special Publication. Volume 4. The University Press of Kansas, Kansas.

Lane, N. G. 1981. A nearshore sponge spicule mat from the Pennsylvanian of west-central Indiana. Journal of Sedimentary Research, 51, 197–202.

Liao, Z. Wang, X. 2002. Fossil ophiuroids from Tongziyan Formation (middle Permian) in Longyan, Fujian. Acta Palaeontologica Sinica, 41, 396–402.

Loriga, C. B. Cavicchi, A. B. 1969. Praeaplocoma hessi n. gen., n. sp: un'Ofiura del Werfeniano (Trias inferiore) del Gruppo della Costabella, Dolomiti. Memorie, Geopaleontologiche, Dell’Universita di Ferrara, 2, 185–197.

Mayou, T. V. 1969. A new species of Permian ophiuroid from Nevada. Journal of Paleontology, 43, 936–940.

Mostler, H. 1971. MikrofaunenausdemUnter-karbonvomhindukusch.Geologisch-PaläontologischeMitteilungen Innsbruck,1(12), 1–19.

Miller, H. W. 1963. Ophiuraster Miller 1958, preoccupied by Ophiuraster Clark 1939. Journal of Paleontology, 37, 725.

Perry, J. A., Glass, A., Kues, B. Ely, L. 2007. Observations on new Late Paleozoic brittle stars (Echinodermata, Ophiuroidea) from New Mexico and Texas [Abstract]. Geological Society of America, Cordilleran Section, 103rd Annual Meeting (4–6 May 2007). Geological Society of America Abstracts with Programs, 39(4), 3.

Posenato, R. 1999. Brachiopods and molluscs. Pp. 104–109 in G. Cassinis, L. Cortesogno, L. Gaggero, F. Massari, C. Neri, U. Nicosia & P. Pittau (eds) Stratigraphy and facies of the Permian deposits between eastern Lombardy and the western Dolomites.The Continental Permian International Congress. Field Trip Guidebook, 23–25, September 1999. Earth Science Department, Pavia University, Pavia.

Runnegar, B. 1969. A lower Triassic ammonoid fauna from southeast Queensland. Journal of Paleontology, 43, 818–828.

Sanchez, T. M. 1983. A new Permian ophiuroid, Archaeophiomusium andinum nov. sp. from western Venezuela. Geobios, 16, 103–107.

Spandel, E. 1898. Die Echinodermen des deutschen Zechsteines. Abhandlungen der Naturhistorischen Gesellschaft zu Nürnberg, 11 for 1897, 17–45.

Spencer, W. K. 1934. Un Ophiuroïde du calcairecarbonifèredu Boulonnais. Annales de la SociétéGéologique du Nord,59, 187–191.

Ward, E. L. 1982. A new ophiuroid from the Pennsylvanian of Kansas, associated with the ichnofossil Asteriacites [Abstract]. Geological Society of America Abstracts with Programs, 14(3), 139.

West, R. R.Ward, E. L. 1990. Asteriacites lumbricalis and a protasterid ophiuroid. Pp. 321–327 in A. J. Boucot (ed.) Evolutionary paleobiology of behavior and coevolution. Elsevier, Amsterdam.

Wu, Q. 1980. First discovery of an upper Permian ophiuroid in Fujian. Acta Palaeontologica Sinica, 19, 61–62.

Yang, T. Y. 1960. On the discovery of a scythic ophiuroid from Kueichou, China. Acta Palaeontologica Sinica, 8, 61–62.

Yang, T. Y., Yin, H. F. Lin, H. M. 1979. Marine Triassic faunas from Shihchienfeng Group in the northern Weihe river basin, Shaanxi Province. Acta Palaeontologica Sinica, 18, 465–474