History of Life from Ordovician

Ordovician Period 488-443 mya

End of the Ordovician was one of the coldest times in Earth history

Marine Diversification

Ordovician - Major diversification of marine organisms: more niches filled!

End of Ordovician

Mass extinction may have been caused by drop in temperature and drop in sea level

Second largest extinction

Silurian Period (444-415 mya)

Gondwana drifts N

Sea level rises and ice sheets melt

Ammonoids

Gnathostomes

Marine vertebrates with jaws and fins

The colonization of land

Plants invaded land in the late Silurian/early Devonian, 408 Ma, probably in wet, transitional habitats

Silurian into Devonian

Devonian Period (416-358 mya)

Pangea starts to form

The colonization of land

Major hurdle: water loss

Key innovations at this time: vascular tissue, support tissue (no water to hold up body); spores, then seeds (with protective covering), for reproduction without water

The colonization of land

Earliest terrestrial animals were arthropods, in Devonian, with marine ancestors

Chelicerates: spiders, scorpions

Mandibulates: millipedes, springtails

Chelicerates

Mandibulates

Herbaceous, predatory, detritivores

Colonization of land

Terrestrial vertebrates arose in late Devonian

Amphibians, fed on arthropods in swampy (transitional) habitats

Although extant tetrapods have five (or fewer) digits, recent discoveries show that early amphibians had 7 or 8 digits

End of Devonian

Mass extinction marked end of Devonian

Carboniferous Period (354-290 mya)

Paleozoic oceans between Euramerica and Gondwana began to close, forming the Appalachian and Variscan mountains

Sea levels swing widely

The first forests

Diversification of seed ferns, gymnosperms, no flowering plants yet

First winged insects, including dragonflies with 70 cm wingspans!

The Amniote Egg

Amniotes include ‘reptiles’, birds, mammals; diverged late Carboniferous

Derived from amphibians

Amniote egg allowed terrestrial reproduction: a key innovation

Amniotes

Anapsida, Synapsida, Diapsida

Refers to skull openings: none, one, or two

Anapsids are turtles

Synapsids are “mammal-like reptiles”

Diapsids gave rise to lizards, archosauromorphs (crocs, dinosaurs)

Synapsids: ancestors to mammals

Permian Period (299-250 mya)

Vast deserts covered western Pangea during the Permian as reptiles spread across the face of the super continent

Permian Period

Insects, amphibians, and early reptiles in swamp forests

Permian extinction

Largest mass extinction (96% of species)

Lowest sea levels in history

Ammonites, stalked echinoderms, brachiopods reduced

Trilobites and many corals extinct

Extinctions on land less pronounced

The Mesozoic Era

Divided into three periods

Triassic

Jurassic

Cretaceous

Major geologic event was the breakup of Pangea

Pangea allowed land animals to migrate from the South Pole to the North Pole

Life began to rediversify after the great Permian extinction and warm-water faunas spread across TethysSea

Period of warm, mild climate

Mesozoic: the age of reptiles

Both marine and terrestrial forms

Land plants dominated by cycads and gymnosperms

Anatomically ‘advanced’ insect orders, including hymenoptera

Triassic Period (251-199 mya)

Seas repopulated after Permian extinction

First dinosaurs and mammals

Marine reptiles: Lepidosaurs

The Amazing Archosaurs

Pterosaurs: evolved true flight

Quetzalcoatlus

Wingspan 12 m

The Amazing Archosaurs

Ornithischia:

Hadrosaurs

Stegosaurs

The Amazing Archosaurs

Saurischia:

Tyrannosaurus rex

Apatosaurus

The Amazing Archosaurs

Birds: living archosaurs

End of Triassic

Mass extinction

Ammonoids and bivalves decimated

Jurassic Period (200-144 mya)

A wide Tethys ocean separated the northern continents from Gondwana

Continents break up by end of period

Jurassic Period

Radiation of the dinosaurs

Mesozoic marine revolution

Crabs

Bony fishes – telosts appear

Gastropods, bivalves and bryozoans rose to dominance

Mammals

Morganudoconts

Cretaceous Period (145-65 mya)

More continental movement

Cretaceous Period

Surviving dinosaurs diversify

Seedless plants and gymnosperms begin to decline

Rise of Flowering Plants

Radiation of the Angiosperms (125 Mya)

Early angiosperms (flowering plants) were non-woody, had a competitive advantage over gymnosperms (conifers, ginkgo)

Sauropods with long necks may have fed on tall conifers in Jurassic

Shorter Hadrosaurs in Cretaceous may have provided selection pressure that favored rapidly growing angiosperms

End of Cretaceous

Mass extinction

Only one surviving lineage of dinosaurs

K-T Asteroid Impact

Asteroid

Present Cenozoic Era

Two Periods

Tertiary Period

Quaternary Period

Seven Epochs

Tertiary Period (65 mya – 5 mya)

More continental movement

The adaptive radiation of the mammals

First appeared in Triassic, placental mammals and marsupials in late Creteceous, big radiation in the Tertiary

First mammals insectivorous or omnivorous; carnivory associated with diversification

Homeothermy, hair, milk = shared traits; most are viviparous

Mammal radiation

Three subclasses:

Prototheria (platypus, echidna: NOT viviparous)

Mammal radiation

Three subclasses:

Metatheria (marsupials)

Mammal radiation

Three subclasses:

Eutheria (placentals)

Mammal radiation

Sinapsids to therapsids to cynodonts to mammals

Mammal radiation

Major changes over time:

Cranial characters, probably for mobility and speed

Jaw characters, probably for subduing prey

All skeletal characters modified from those existing in ancestors

Mammal radiation

Phylogenetic relationships were difficult to resolve: radiation was fast

Mammal radiation

Angiosperm coevolution with insects

The flower as a key innovation

Synchronicity between adaptive radiation of hymenoptera and lepidoptera, and adaptive radiation of angiosperms, in Tertiary

Insect pollination more efficient than wind pollination

Angiosperm coevolution with insects

Coevolution between plants and insect herbivores also important

Resin canals, a defense, has evolved multiple times

Oligocene

Adaptive radiation of Poaceae

Quaternary Period (5 mya-present)

•The world has taken on a "modern" look, but note that Florida and parts of Asia were flooded by the sea

Pleistocene

The last expansion of the polar ice sheets took place about 18,000 years ago

Ice age

Sea levels dropped

70% of mollusc species became extinct

Global climate generally drier

Later Cenozoic

Climates became cooler and drier

Grasslands and woodlands dominated

Grazing and browsing animals thrived

Earth in ice age

Extinctions in Pleistocene

Shallow water marine invertebrates

Large bodied mammals and birds

Ground sloth

Giant bison

Saber tooth tiger

Current extinction

We are perhaps in the midst of another mass extinction

Earth today