Explanations for Plant and Animal Distributions

I. Ecological

A. Tolerance / Limits

Bioclimatic Frontiers

B. Species Interactions

C. Succession

Primary, Secondary

II. Historical

A. Evolution

B. Speciation

C. Extinction

D. Dispersal

I. Ecological Biogeography

A. Tolerance / Limit

Plant’s survival depends on tolerance of environment

Climate, soil, biota, topography, fire

evolved means of coping with various environments: adaptations

e.g.: drought

Examples of Types of Adaptations:

Xerophytic: adapted to drought

Examples:

1. Waxy coatings on leaves (seal in water vapor)

Tropophytic: adapted to wet-dry (heavy rain/ drought) climates

Example: Deciduous trees: drop leaves after wet season; dormant during dry

Hydrophytic: water-tolerant; require saturation

Halophytic: salt-tolerant

Animal adaptationsto temperature:

Cold-blooded: no internal temperature regulation

Active during warm; dormant during cold

Hibernation

Warm-blooded: metabolism maintains constant temperature

Fur, hair, feathers, fat

Sweating, panting

Bioclimatic frontiers

Boundary marking limits of species’ tolerance to environmental factors

B. Species Interactions

1) Competition: both species require a common resource; niche overlap

And NUH is the letter I use to spell Nutches,Who live in small caves, known as Niches, for hutches.These Nutches have troubles, the biggest of which isThe fact there are many more Nutches than Niches.Each Nutch in a Nich knows that some other NutchWould like to move into his Nich very much.So each Nutch in a Nich has to watch that small NichOr Nutches who haven't got Niches will snitch.

Dr. Seuss - On Beyond Zebra (1955)

2) Predation: one species preys on (eats) another

3) Parasitism: one species gains nutrition from the other

4) Symbiosis: some benefit gained

Commensalism: one benefits, other not affected

Mutualism: both benefit

C. Ecological Succession

Change in species composition, structure or architecture of vegetation through time.

Sequence of communities that blend into one another gradually over time.

Development of plant community is gradual !

various communities occupy a habitat until a final, stable CLIMAX community is established.

May take <100 – 1000’s years

Several different sequences can produce same climax community

Example: Isle Royale….

Primary vs. Secondary Succession:

Primary: occurs on sites with no (or very little)vestige of a former community

Mt. St. Helens
Cooled lava flow
Strip mines

Secondary: habitat not completely sterile; some members of previous community present

Forest fire (mild)
Abandoned agricultural field
Flooded land

Examples of succession:

1. Pond Bog Forest

2. New volcanic eruption

Krakatau example:

9 months: tiny spider was found

>12 months: grasses

< 50 years: dense rain forest

How did seeds get there?

Wind, carried on/in animals

Swimming, rafting

storms

3. Old field

About succession… Climax community depends on climate

Colonization of bare ground depends on availability of seeds

Same climax community can be reached through different series of steps

Succession modifies environment

II. Historical biogeography

A. Evolutionexplains the history and biodiversityof life on Earth

40,000 microorganisms

350,000 plants

2.2 million animals

800,000 insects (1/3)

Definition: descent with modification:

All life shares common ancestor.

By having generation after generation of descendants with modifications, the immense diversity of life arose.

Key is variation:Difference between parent and offspring

Darwin 1859 :

Variationacted upon by natural selection over time brings about new species.

Favorable traits become more common in successive generations of a population. Variations are inherited; those that survive are those with variations that enhance their survival.

Darwin lacked mechanism for variation

Mechanisms of evolution:mutation and recombination

Has to be a change in genetic frequency in a population over time.

Mutations
altered genetic material; create genetic variation
Most are benign or not beneficial; some increase survival rate (those are passed on)
Recombinations
Each offspring gets 2 different alleles (copies) of each gene; many 1000s of genes per organism yields enormous possible genetic combinations : variations

B. Speciation

Means by which species are differentiated and maintained:

Mutation

Natural selection

Genetic Drift: change in gene pool due to chance rather than natural selection

Geographic Isolation : populations diverge and lose ability to interbreed

C. Extinction

Conditions change more rapidly than a population’s ability to evolve new adaptations.

D. Dispersal

Species move from one location to another; colonize new areas

Example: hemlock exercise in lab