American Museum of Natural History

David A. Grimaldi November, 2015

Division of Invertebrate Zoology

American Museum of Natural History

Central Park West at 79th Street

New York, New York 10024-5192

Telephone: 212-769-5615, -5277 FAX

Email:

RESEARCH AREAS

Diversity, fossil record, and evolution of insects; fossilization in amber; systematics of Drosophilidae and other Diptera; evolution of sociality in insects; the evolution of biological communities, particularly tropical forests; extinction and environmental deterioration.

EDUCATION

B. Sc., 1979, University of Connecticut. Major concentrations in Biology and Fine Arts.

M.Sc., 1983, State University of New York, Binghamton. Evolutionary Biology.

Ph.D., 1986, Cornell University, Entomology.

POSITIONS

1986-1991: Assistant Curator, American Museum of Natural History.

1988-present: Adjunct Professor, City University of New York (Graduate Program in Ecology and Evolutionary Biology).

1991-1996: Associate Curator (tenured), American Museum of Natural History.

1994-present: Adjunct Professor, Cornell University (Department of Entomology).

1994-1999: Chairman, Department of Entomology, American Museum of Natural History.

1996-present: Curator, American Museum of Natural History.

1995-present: Adjunct Professor, Columbia University (Department of Ecology, Evolution, and Environmental Biology).

2008-present: Professor, Gilder Graduate School, Program in Comparative Biology, American Museum of Natural History

AWARDS

________________________________________________________________________

2007: Thomas Say Award, Entomological Society of America, for distinguished contributions to insect systematics.

2015: Gilbert Harris Award, Paleontological Research Institute, for distinguished contributions to systematic paleontology.

GRANTS AND FUNDING: last 15 years

1998-2001: NSF, Biological Infrastructure Program, $90,000, for a databased catalogue of the world species of termites (Isoptera). PI: Kumar Krishna, co-PI: D. Grimaldi.

2000-2002: NSF, Biological Collections Program, $292,000, for curation, conservation, and databasing of the collections of fossil insects and amber fossils at the AMNH. PI: D. Grimaldi

2000-2003: NSF, Systematic Biology Program, $245,000, for a monographic revision of the Drosophila melanogaster species group. PI: D. Grimaldi; co-PI: Valerie Schawaroch.

2001-2004: NSF, Geosciences Program, $240,000, for research on Triassic fossils from the Cow Branch Formation of Virginia. PI: Nick Fraser. Co-PI: David Grimaldi

2006-2009: NSF, Systematics Program, $116,000, for research on insects in Cretaceous amber and the origin of modern biological communities. PI: David Grimaldi

2011-2013: Institute for Museum and Library Services, $149,000, for conservation of fossiliferous amber specimens in the AMNH. PI: Lisa Elkin, Co-PI: D. Grimaldi.

2012-2013: NSF, DDIG, $14,500: A time scale for scales: Phylogeny and divergence times of basal lineages of Coccoidea (Insecta: Hemiptera). CoPI: Isabelle Vea (RGGS Student).

2013-2014: NSF DDIG, $15,527: Phylogeny and fossil record of the ants. CoPI: Phil Barden (RGGS student).

2013-2015: National Science Foundation EF-1304943, “Digitization TCN: Collaborative Research: Fossil Insect Collaborative: A deep-time approach to studying diversification and response to environmental change.” A collaborative grant involving 7 institutions; AMNH award is $352,484.

2016-2018: National Science Foundation OPUS Award, "The Brachyceran Flies: Synthesizing 30 Years of Research on 200 Million Years of Evolutionary History." PI: D. Grimaldi.

TEACHING AND MENTORING*: last 10 years

UNDERGRADUATES (*NSF Research Experience for Undergraduates program):

James Hayden, Columbia Univ., 2003: Diversity and community ecology of Macrolepidoptera.

Ann Carla Staver, Columbia Univ., 2004: Diversity and abundance in forest canopy Drosophilidae

Megan Adcock, Bennington College, 2004: Systematics of fossil and living emesine bugs.

Aaron Davitt, University of Denver, 2005: Mesozoic Coleoptera.

Sweta Agrawal, Columbia University, 2008: Comparative study of the halteres in Diptera.

Phillip Barden, Arizona State University, 2009: Comparative morphology of the termite fontanelle

GRADUATE STUDENT COMMITTEES/MENTORING: (Major Advisor): last 10 years

Caroline Chaboo, Ph.D., 2005, Cornell Univ.: Systematics of cassidine Chrysomelidae (tortoise beetles). Presently: Assistant Professor, University of Kansas.

Sara Lubkin, Ph.D., 2007, Cornell Univ: Systematics and paleontology of Paleozoic and Mesozoic Archostemata (Coleoptera).

Craig Gibbs, Ph.D., CUNY: Patterns of drosophilid fly species diversity and abundance in Neotropical forest canopies. Presently: Queens Zoo, WCS.

Torsten Dikow, Ph. D., 2007, Cornell Univ.: Phylogeny of the robber flies (Diptera: Asilidae). Presently: Research Scientist, Field Museum (Chicago).

Isabelle Vea, currently (2009-2013): Phylogeny and fossil record of the archaeococcoid scale insects. Presently: Postdoctoral researcher, Univ. Nagoya, Japan.

Philip Barden, currently (started 2010): Phylogeny and fossil record of the basal lineages of ants.

POSTDOCTORAL FELLOWS: last 10 years

Valerie Schawaroch, 2000-2004, AMNH: A monographic revision of the Drosophila melanogaster species group (NSF Fellow). Presently: Associate Professor, Baruch College, CUNY.

Enrique Penalver, 2004-2006, Univ. of Valencia, Spain: Paleobiology and paleoecology of the Miocene fossil community preserved in Dominican amber (Spanish government funded). Presently: Curator, Museo Geominero, Madrid.

Jessica Ware, 2008-2010: Species revision and phylogeny of the lower termites (NSF Fellow). Presently: Assistant Professor, Rutgers Univ.

Steven Ray Davis, 2014-2016: Transcriptome studies on the diversity and evolution of weevil rostrum (Gerstner Postdoctoral Fellow).

COURSES TAUGHT: last 10 years

Columbia Univ., Insect Diversity (2001, 2003, 2005, 2007, 2014) .

COLLECTION STEWARDSHIP

Curator-in-Charge of Diptera, Lepidoptera, Isoptera, fossil insects, and minor holometabolan orders (Trichoptera, Mecoptera, Siphonaptera, Strepsiptera).

Development of world's largest collection of amber fossils, through acquisitions and fieldwork, including 8,000 specimens of amber fossils from the Miocene of the Dominican Republic and Oligocene of Mexico; 3,000 specimens of Eocene amber from Baltic region and India; and 7,000 specimens from the Cretaceous of New Jersey, Lebanon, Burma (Myanmar). Studies on conservation, preparation, imaging, and treatment of amber.

Development of largest and most diverse collection of Drosophilidae (Diptera) in the world, c. 400,000 specimens and 2,000 species, largely through fieldwork.

INVITED LECUTRES: last 7 years.

35 invited lectures were given at various universities and conferences, in Asia, Australia, Europe, North America, and Africa.

TRAVEL (MUSEUMS, ETC.) AND FIELDWORK

AFRICA:

Tanzania, 1995, biodiversity survey of Tarangire National Park.

Republic of South Africa, 2005: Kruger National Park; Pretoria (work at National Botanical Inst.)

ASIA:

Vietnam, 1998-2000, biodiversity surveys of protected forests (sponsored by NSF).

India, 2008-10, 2013, 2015: excavations of Paleo-Eocene amber from Gujarat, western India.

EUROPE:

Austria, 1996: Vienna, Salzburg

Denmark, 1998, work at Zoological Museum, Univ. Copenhagen.

England, 1990, 1995, 1999, 2000, 2002, work at Natural History Museum, London; Oxford U.

Germany, 1994; 1996, 2009, 2010, 2011: work at Stuttgart Museum; Humboldt Museum, Berlin.

Greece, 2007: Athens, Crete

Hungary, 1996, 2011: Budapest (work at Hungarian Natural History Museum).

Italy, 1995-2000: work at museums in Florence, Milan, Turin, Venice.

Poland: Pongracz, Cracow, Warsaw.

Russia, 1995, St. Petersburg, work at Zool. Instit., Ekaterininsky Palace; Moscow (Paleo. Inst.) 2002.

Slovakia, 1992, Bratislava: meetings.

The Netherlands, 1995: work at Leiden museum.

PACIFIC:

Hawaii, 1993: fieldwork on all the large islands

Fiji Islands, 2008: fieldwork on Vanua Levu and Viti Levu.

New Caledonia, 2008: fieldwork.

NORTH AMERICA:

Alaska, 1998, 2001: prospecting Cretaceous on peninsula, Kodiak Is., arctic tundra

Arizona, 1990-2000: field work on Diptera in southeastern Arizona.

Arkansas, 1992: field work on Diptera; excavations of Eocene amber (Claiborne Form.)

Florida, 1995-2000: field work on Diptera, principally in Everglades.

New Jersey, 1992-1998: excavations of Cretaceous amber.

South Dakota, 2010: Field work in Hell Creek Formation, Harding County.

Virginia: 1998-present: excavations of Triassic insects from the Cow Branch Formation.

Wyoming, 1997-1999: prospecting and excavations of K/T amber.

Washington state, 1994: Excavations of Eocene amber from Renton Formation.

Washington, D.C., 1985-present: yearly work at Smithsonian Institution.

Eastern United States, 1990- present: prospecting and excavations for Cretaceous amber

outcrops.

SOUTH AND CENTRAL AMERICA, CARIBBEAN:

Costa Rica, 1989, 1991, 1993, 1995, 1999, 2013: Field work on Diptera.

Brazil, 2000: Field work on Diptera, plenary lecture at Int. Congr. Ent.

Dominican Republic, 1990, 1991, 1992, 1995, 2014: Biodiversity surveys of modern insect

fauna and stratigraphic work on Miocene amber deposits.

French Guiana, 1997: Field work on Diptera.

Panama, 1985, 1988, 1999: Field work on Diptera.

Peru, 1984: Field work on Diptera.

Venezuela, 1989, 1991, 1993: Biodiversity surveys of tepuis; field work in Andes.

Virgin Islands (British and U.S.), 1990, 1992, 1993: Biodiversity surveys.

PUBLICATIONS

1. Jaenike, J., S.Ausubel, and D.A. Grimaldi. 1982. On the evolution of clonal diversity in parthenogenetic earthworms. Pedobiologia 23: 304-310.

2. Grimaldi, D.A., and J. Jaenike. 1982. Laboratory culturing of mycophagous drosophilids. Dros. Info. Serv. 58: 156-157.

3. Grimaldi, D. and J. Jaenike. 1983. The Diptera breeding on skunk cabbage, Symplocarpus foetidus (Araceae). J. New York Ent. Soc. 91: 83-89.

4. Jaenike, J. and D. Grimaldi. 1983. Genetic variation for host preference within and among populations of Drosophila tripunctata. Evolution 37: 1023-1033.

5. Jaenike, J., D.A. Grimaldi, A.E. Sluder, and A.L. Greenleaf. 1983. a-amanitin tolerance in mycophagous Drosophila. Science 221: 165-167.

6. Grimaldi, D. 1984. Review of: Biological Diversification in the Tropics: Proceedings of the 5th International Symposium for the Association for Tropical Biology, G.T. Prance (ed.), Columbia Univ. Press, 714 pp. J. New York Ent. Soc. 92: 94-96.

7. Grimaldi, D. and J. Jaenike. 1984. Competition in natural populations of mycophagous Drosophila. Ecology 65: 1113-1120.

8. Grimaldi, D. 1985. Niche separation and competitive coexistence in mycophagous Drosophila (Diptera: Drosophilidae). Proc. Ent. Soc. Washington 87: 498-511.

9. Grimaldi, D. 1985. Review of: Courtship Behaviors of the Hawaiian Picture-Winged Drosophila, by H.T. Spieth, 1984, Univ. Calif. Publ. Entomol. 103 (92 pp.). J.New York Entomol. Soc. 1285-1287.

10. Grimaldi, D. 1986. The Chymomyza aldrichii species-group (Diptera: Drosophilidae): relationships, new Neotropical species, and the evolution of some sexual traits. J. New York Ent. Soc. 94: 342-371.

11. Grimaldi, D. 1986. A new Drosophila (Hirtodrosophila) from Malaysia with broad-headed males (Diptera: Drosophilidae). J. New York Ent. Soc. 94: 372-376.

12. Grimaldi, D. and B.A. Underwood. 1986. Megabraula, a new genus for two new species of Braulidae (Diptera), and a discussion of braulid evolution. Syst. Entomol. 11: 427-438.

13. Grimaldi, D.A. 1987. Phylogenetics and taxonomy of Zygothrica (Diptera: Drosophilidae). Bull. Amer. Mus. Nat. Hist. 186: 103-268.

14. Grimaldi, D.A. 1987. Amber fossil Drosophilidae (Diptera), with particular reference to the hispaniolan taxa. Amer. Mus. Novitates 2880: 23 pp.

15. Grimaldi, D.A. 1988. Relicts in the Drosophilidae (Diptera). Pp. 183-213, In: J.K. Liebherr (ed.), Zoogeography of Caribbean Insects. Ithaca: Cornell Univ. Press.

16. Grimaldi, D. 1988. Bee flies and bluets: Bombylius (Diptera: Bombyliidae) flower-constant on the distylous species, Hedyotis caerulea (Rubiaceae), and the manner of foraging. J. Nat. Hist. 22: 1-10.

17. Grimaldi, D. 1988. Review of: Manual of Nearctic Diptera, vol. 2. J.F. McAlpine (ed.) Research Branch Agriculture Canada Monogr. 28: 657 pp. J. New York Ent. Soc. 96: 124-126.

18. Grimaldi, D. 1988. Drosophila (Hirtodrosophila) chandleri (Diptera: Drosophilidae) a new species from Sri Lanka with broad-headed males. J. New York Ent. Soc. 96: 323-326.

19. Michener, C.D. and D.A. Grimaldi. 1988. A Trigona from Late Cretaceous amber of New Jersey (Hymenoptera: Apidae: Meliponinae). Amer.Mus. Novitates 2917: 10 pp.

20. Michener, C.D. and D. Grimaldi. 1988. The oldest fossil bee: apoid history, evolutionary stasis, and antiquity of social behavior. Proc. Nat. Acad. Sci. (USA) 85: 6424-6426.

21. Grimaldi, D. 1988. Still life with flowers: the discovery of the oldest fossil bee. Natural History 9/88: 86-89.

22. Grimaldi, D. 1989. The genus Metopina (Diptera: Phoridae) from Cretaceous and Tertiary ambers. J. New York Ent. Soc. 97: 65-72.

23. Grimaldi, D. and G. Fenster. 1989. Evolution of extreme sexual dimorphisms: structural and behavioral convergence among broad-headed male Drosophilidae (Diptera). Amer. Mus. Novitates 2939: 25 pp.

24. Grimaldi, D., C.W. Beck, and J.J. Boon. 1989. Occurrence, chemical characteristics, and paleontology of the fossil resins from New Jersey. Amer. Mus. Novitates 2948: 28 pp.

25. Grimaldi, D. (ed.) 1990. Insects from the Santana Formation, Lower Cretaceous, of Brazil. Bull. Amer. Mus. Nat. Hist. 195: 191 pp.

26. Grimaldi, D. 1990. Diptera, pp. 164-183, In: ibid.

27. Grimaldi, D. 1990. New distributional records of Scaptomyza (Bunostoma) australis from South Pacific islands and biogeographic implications. J. New York Ent. Soc. 98: 484-488.

28. Grimaldi, D. 1990. A phylogenetic classification of the Drosophilidae. Pg. 344, In: Abstr. 2nd Int. Congress Dipterology, Bratislava, Aug. 27-Sept. 1, 1990.

29. Grimaldi, D. 1990. A phylogenetic, revised classification of genera in the Drosophilidae (Diptera). Bull. Amer. Mus. Nat. Hist. 197: 139 pp.

30. Grimaldi, D. 1990. Revision of Zygothrica (Diptera: Drosophilidae), Part II. The first African species, two new Indo-Pacific groups, and the bilineata and samoaensis species groups. Amer. Mus. Novitates 2964: 31 pp.

31. Poinar, G.Q., Jr. and D.A. Grimaldi. 1990. Fossil and extant macrochelid mites (Acari: Macrochelidae) phoretic on drosophilid flies (Diptera: Drosophilidae). J. New York Entomol. Soc. 98: 88-92.

32. Grimaldi, D. 1991. Mycetobiine woodgnats (Diptera: Anisopodidae) from the Oligo-Miocene amber of the Dominican Republic, and Old World affinities. Amer.Mus.Novitates 3014: 24 pp.

33. Grimaldi, D. 1991. Drosophila canaryana Takada and Yoon, 1989 (Diptera: Drosophilidae), a junior synonym of Drosophila guanche Monclús, 1976. Entomol. News 102: 223-226.

34. Grimaldi, D. 1991. Cladistics and the classification of the Drosophilidae. Pp. 85-97, In: L. Weismann et al. (eds.), Proc. 2nd Int. Congr. Dipterology (Bratislava). SPB Acad. Publ.: The Hague.

35. Grimaldi, D. 1991. The Santana Formation insects. Pp. 379-406, In: J.G. Maisey (ed.), Santana Fossils: An Illustrated Atlas. New Jersey: T.F.H.Publications.

36. DeSalle, R. and D.A. Grimaldi. 1991. Morphological and molecular systematics of the Drosophilidae. Annu. Rev. Ecol. Syst. 22: 447-475.

37. Heed, W.B. and D.A. Grimaldi. 1991. Revision of the morphocryptic, Caribbean mayaguana species subcluster in the Drosophila repleta group (Diptera: Drosophilidae). Amer. Mus. Novitates 2999: 15 pp.

38. Krishna, K. and D. Grimaldi. 1991. A new fossil species from Dominican amber of the living Australian termite genus Mastotermes (Isoptera: Mastotermitidae). Amer. Mus. Novitates 3021: 10 pp.

39. Shedrinsky, A.M., D. Grimaldi, T.P. Wampler, and N.S. Baer. 1991. Amber and copal: pyrolysis gas chromatographic (PyGC) studies of provenance. Wien. Ber. Naturwiss. Kunst 6/7/8: 37-62.

40. Grimaldi, D. 1992. Systematics of the genus Colocasiomyia de Meijere (Diptera: Drosophilidae): cladistics, a new generic synonym, new records, and a new species from Nepal. Entomol. Scandanavica 22: 417-426.

41. Grimaldi, D. 1992. Drosophila putrida Sturtevant, 1916 (Insecta, Diptera): proposed replacement of the holotype by a neotype. Bull. Zool. Nomenclature 49: 129-132.

42. Grimaldi, D. and C. Young. 1992. Observations on the bizarre jelly mass habit of larval Geranomyia (Diptera: Tipulidae: Limoniidae). J. New York Ent. Soc. 100: 634-637.

43. Grimaldi, D. et al. 1992. Systematics and modes of reproductive isolation in the Holarctic Drosophila testacea species group (Diptera: Drosophilidae). Ann. Entomol. Soc.Amer. 85: 671-685.

44. Grimaldi, D. 1992. Vicariance biogeography, geographic extinctions, and the North American Oligocene tsetse flies. Pp. 178-204, In: M.J. Novacek and Q.D. Wheeler (eds.), Extinction and Phylogeny. New York: Columbia Univ. Press.

45. DeSalle, R. and D. Grimaldi. 1992. Characters and the systematics of the Drosophilidae. J. Heredity 83: 182-188.

46. DeSalle, R., J. Gatesy, W. Wheeler,and D. Grimaldi. 1992. DNA sequences from a fossil termite in Oligo-Miocene amber and their phylogenetic implications. Science 257: 1933-1936.