Appendix 1 Proboscidean taxa and published data sources used in the comparative study of the Woranso-Mille proboscidean sample. More complete lists of site samples for each taxon are provided in Sanders et al. (2010).
Taxon / Site / Original Specimens Studied? / Published Data
Proboscidea
Gomphotheriidae
Anancus kenyensis / Lothagam, Kenya / yes / Tassy 2003
Mpesida Beds, Kenya / yes / Tassy 1986
Middle Awash, Ethiopia / yes / Mebrate and Kalb 1985; Kalb and Mebrate 1993; Kalb and Froehlich 1995
Lemudong’o, Kenya / Saegusa and Hlusko 2007
Manonga Valley, Tanzania / yes / Sanders 1997
Kanam East and West, Kenya / yes / MacInnes 1942
Nkondo, Uganda / yes / Tassy 1995
Gona, Ethiopia / yes / Semaw et al. 2005
Endolele, Tanzania / yes / Sanders 2011
Galili, Ethiopia / Kullmer et al. 2008
Anancus ultimus / Tugen Hills, Kenya / yes / Hill et al. 1985, 1986
Lothagam, Kenya / yes / Tassy 2003
Nyawiega, Uganda / yes / Cooke and Coryndon 1970; Tassy 1995
Manonga Valley, Tanzania / yes / Sanders 1997
Middle Awash, Ethiopia / yes / Mebrate and Kalb 1985; Kalb and Mebrate 1993; Kalb and Froehlich 1995
Kakesio, Tanzania / yes / Sanders 2011
Kanapoi, Kenya / yes / Harris et al. 2003
Laetoli, Tanzania / yes / Sanders 2011
Elephantidae
Stegotetrabelodon syrticus / Sahabi, Libya / Petrocchi 1943, 1954; Maglio 1973; Sanders 2008
Stegotetrabelodon orbus / Lothagam, Kenya / yes / Maglio 1970b, 1973; Maglio and Ricca 1977; Tassy 2003
Tugen Hills, Kenya / yes / Hill et al. 1985, 1986; Tassy 1986
Manonga Valley / yes / Sanders 1997
Stegodibelodon schneideri / Menalla, Chad / Coppens 1972; Coppens et al. 1978
Kollé, Chad / Mackaye et al. 2005; Sanders et al. 2010
Primelephas korotorensis / Lothagam, Kenya / yes / Maglio 1970b, 1973; Maglio and Ricca 1977; Tassy 2003
Toros Menalla, Chad / Mackaye et al. 2008
Tugen Hills, Kenya / yes / Hill et al. 1985, 1986; Tassy 1986
Middle Awash, Ethiopia / yes / Mebrate 1983; Kalb and Mebrate 1993; Kalb 1995; Haile-Selassie 2001; Saegusa and Haile-Selassie 2009
Manonga Valley / yes / Sanders 1997
Nyabusosi and North and South Nyabrogo, Uganda / yes / Cooke and Coryndon 1970; Tassy 1995
Kossom Bougoudi and Kollé, Chad / Mackaye et al. 2008
Koulà and Kolinga, Chad / Coppens 1965, 1967; Maglio 1973; Coppens et al. 1978
Mammuthus subplanifrons / Middle Awash, Ethiopia / yes / Mebrate 1983; Kalb and Mebrate 1993; Haile-Selassie 2001
Langebaanweg, South Africa / yes / Maglio and Hendey 1970; Sanders 2006 2007
Mammuthus sp. / Middle Awash, Ethiopia / Saegusa and Haile-Selassie 2009
Mammuthus sp. “Hadar type” / Middle Awash, Ethiopia / yes / Beden 1979, 1985; White et al. 1984; Kalb and Froehlich 1995
Mammuthus africanavus / Lac Ichkeul, Tunisia / yes / Arambourg 1970; Maglio 1973; Sanders et al. 2010; Markov in press
Garet et Tir, Algeria / Arambourg 1970, Sanders et al. 2010; Markov in press
Aïn Brimba, Tunisia / Arambourg 1970, Sanders et al. 2010; Markov in press
Loxodonta cookei / Nkondo-Kaiso, Uganda / yes / Tassy 1995; Sanders 2007
Langebaanweg, South Africa / yes / Sanders 2006, 2007
Endolele, Tanzania / yes / Sanders 2011
Loxdonta exoptata / Kakesio, Tanzania / yes / Sanders 2011
Kanapoi, Kenya / yes / Harris et al. 2003
Allia Bay, Kenya / yes / Beden 1983
Laetoli, Tanzania / yes / Beden 1987b; Sanders 2011
West Turkana, Kenya / Harris et al. 1988
Omo, Ethiopia / Beden 1987a
Hadar, Ethiopia / yes / White et al. 1984; Kalb and Mebrate 1993
Loxodonta adaurora adaurora / Kanapoi, Kenya / yes / Maglio 1970b, 1973; Harris et al. 2003
Omo, Ethiopia / Beden 1987a
West Turkana, Kenya / Harris et al. 1988
Allia Bay, Kenya / yes / Beden 1983
Elephas ekorensis / Kanapoi, Kenya / yes / Harris et al. 2003
Ekora, Kenya / Maglio 1970b, 1973
Hadar, Ethiopia / yes / White et al. 1984
Omo, Ethiopia / Beden 1987a
Elephas recki brumpti / Koobi Fora and Allia Bay, Kenya / yes / Beden 1983
Omo, Ethiopia / Beden 1980, 1987a
West Turkana, Kenya / Harris et al. 1988
Middle Awash, Ethiopia / yes / Kalb and Mebrate 1993
Elephas recki shungurensis / Koobi Fora and Allia Bay, Kenya / yes / Beden 1983
Middle Awash, Ethiopia / yes / White et al. 1984
West Turkana, Kenya / Harris et al. 1988
Omo, Ethiopia / Beden 1980, 1987a
Elephas recki atavus / Omo, Ethiopia / Beden 1980, 1987a
Koobi Fora and Ileret, Kenya / yes / Beden 1983
Olduvai Gorge, Tanzania / Beden 1985
Elephas recki ileretensis / Omo, Ethiopia / Beden 1980, 1987a
Koobi Fora and Ileret, Kenya / yes / Beden 1983
Olduvai Gorge, Tanzania / Beden 1985
Elephas recki recki / Koobi Fora and Ileret, Kenya / yes / Beden 1983
Omo, Ethiopia / Beden 1987a
West Turkana, Kenya / Harris et al. 1988
Olduvai Gorge, Tanzania / Beden 1985

Arambourg C (1970) Les Vertébrés du Pléistocène de l’Afrique du Nord. Arch Mus Natl His Nat Paris 7e Série 10:1-126

Beden M (1979) Données récentes sur l’évolution des Proboscidiens pendant le Plio-Pléistocène en Afrique Orientale. Bull Soc Geol France 21:271-276

Beden M (1980) Elephas recki Dietrich, 1915 (Proboscidea, Elephantidae), Évolution au cours du Plio-Pléistocène en Afrique orientale. Geobios 13:891-901

Beden M (1983) Family Elephantidae. In: Harris JM (ed) Koobi Fora Research Project, Vol. 2. Clarendon Press, Oxford, pp 40-129

Beden M (1985) Les Proboscidiens des grands gisements à hominidés Plio-Pléistocènes d’Afrique Orientale. In: Beden M, Behrensmeyer AK, Boaz NT, Bonnefille R, Brain CK, Cooke B, Coppens Y, Dechamps R, Eisenmann V, Gentry A, Geraads D, Gèze R, Guérin C, Harris J, Koeniguer J-C, Letouzey R, Petter G, Vincens A, Vrba E (eds) L’Environnement des Hominides au Plio-Pleistocene. Masson, Paris, pp 21-44

Beden M (1987a) Les Faunes Plio-Pléistocènes de la Vallée de l’Omo (Éthiopie). Tome 2. Les Elephantidés (Mammalia, Proboscidea). Éditions du Centre National de la Recherche Scientifique, Paris

Beden M (1987b) Fossil Elephantidae from Laetoli. In: Leakey MD, Harris JM (eds) Laetoli: A Pliocene Site in Northern Tanzania. Clarendon Press, Oxford, pp 259-294

Cooke HBS, Coryndon S (1970) Fossil mammals from the Kaiso Formation and other related deposits in Uganda. In: Leakey LSB, Savage RGJ (eds) Fossil Vertebrates of Africa, Vol. 2. Academic Press, Edinburgh, pp 107-224

Coppens Y (1965) Les Proboscidiens du Tchad. Actes du Ve Congrés Panafricain de Préhistoire et de l’Étude du Quaternaire (Santa Cruz de Tenerife), 1, 5:331-387

Coppens Y (1967) Les Faunes de Vertebres Quaternaires du Tchad. In: Bishop WW, Clark JD (eds) Background to Evolution in Africa. University of Chicago Press, Chicago, pp 89-97

Coppens Y (1972) Un nouveau Proboscidien du Pliocène du Tchad, Stegodibelodon schneideri nov. gen. nov. sp., et le phylum Stegotetrabelodontinae. C R Acad Sci D, 274:2962-2965

Coppens Y, Maglio VJ, Madden CT, Beden M (1978) Proboscidea. In: Maglio VJ, Cooke HBS (eds) Evolution of African Mammals. Harvard University Press, Cambridge, pp 336-367

Haile-Selassie Y (2001) Late Miocene mammalian fauna from the Middle Awash Valley, Ethiopia. Unpublished PhD dissertation, University of California, Berkeley

Harris JM, Brown FH, Leakey MG (1988) Geology and palaeontology of Plio-Pleistocene localities west of Lake Turkana, Kenya. Contrib Sci Nat Hist Mus LA County 399:1-128

Harris JM, Leakey MG, Cerling TE (2003) Early Pliocene tetrapod remains from Kanapoi, Lake Turkana Basin, Kenya. Contrib Sci Nat Hist Mus LA County 498:39-113

Hill A, Curtis G, Drake R (1986) Sedimentary stratigraphy of the Tugen Hills, Baringo, Kenya. In: Frostick LE, Renaut RW, Reid I, Tiercelin J-J (eds) Sedimentation in the African Rifts. Blackwell and Geological Society of London Special Publication 25, Oxford, pp 285-295

Hill A, Drake R, Tauxe L, Monaghan M, Barry JC, Behrensmeyer AK, Curtis G, Jacobs BF, Jacobs L, Johnson N, Pilbeam D (1985) Neogene palaeontology and geochronology of the Baringo Basin, Kenya. J Hum Evol 14:759-773

Kalb JE (1995) Fossil elephantoids, Awash paleolake basins, and the Afar triple junction, Ethiopia. Palaeogeogr Palaeoclimat Palaeoecol 114:357-368

Kalb JE, Froehlich DJ (1995) Interrelationships of late Neogene elephantoids: new evidence from the Middle Awash Valley, Ethiopia. Geobios 28:727-736

Kalb JE, Mebrate A (1993) Fossil elephantoids from the hominid-bearing Awash Group, Middle Awash Valley, Afar Depression, Ethiopia. Trans Am Philos Soc 83:1-114

Kullmer O, Sandrock O, Viola TB, Hujer W, Said H, Seidler H (2008) Suids, elephantoids, paleochronology, and paleoecology of the Pliocene hominid site Galili, Somali Region, Ethiopia. Palaios 23:452-464

MacInnes DG (1942) Miocene and post-Miocene Proboscidia [sic] from East Africa. Trans Zool Soc Lond 25:33-106

Mackaye HT, Brunet M, Tassy P (2005) Selenetherium kolleensis nov. gen. nov. sp.: un nouveau Proboscidea (Mammalia) dans le Pliocène tchadien. Geobios 38:765-777

Mackaye HT, Coppens Y, Vignaud P, Lihoreau F, Brunet M (2008) De nouveaux restes de Primelephas dans le Mio-Pliocène du Nord du Tchad et revision du genre Primelephas. C R Palevol 7:227-236

Maglio VJ (1970) Four new species of Elephantidae from the Plio-Pleistocene of northwestern Kenya. Breviora 341:1-43

Maglio VJ (1973) Origin and evolution of the Elephantidae. Trans Am Philos Soc 63:1-149

Maglio VJ, Hendey QB (1970) New evidence relating to the supposed stegolophodont ancestry of the Elephantidae. S Afr Archaeol Bull 25:85-87

Maglio VJ, Ricca AB (1977) Dental and skeletal morphology of the earliest elephants. Verh K Ned Akad Wet Afd Natuurkd Erste Reeks 29:1-51

Markov GN (in press) Mammuthus rumanus, early mammoths, and migration out of Africa: some interrelated problems. Quaternary Internatl

Mebrate A (1983) Late Miocene-middle Pleistocene proboscidean fossil remains from the Middle Awash Valley, Afar Depression, Ethiopia. Unpublished Master’s Thesis, Department of Systematics and Ecology, University of Kansas

Mebrate A, Kalb JE (1985) Anancinae (Proboscidea: Gomphotheriidae) from the Middle Awash Valley, Afar, Ethiopia. J Vertebr Paleontol 5:93-102

Petrocchi C (1943) Il giacimento fossilifero di Sahabi. Collezione scientifica e documentaria del Ministero dell’ Africa Italiana, Verbania

Petrocchi C (1954) Paleontologia di Sahabi. Parte I. Proboscidati di Sahabi. Rend Accad Naz XL 4-5:8-74

Saegusa H, Haile-Selassie Y (2009) Proboscidea. In: Haile-Selassie Y, WoldeGabriel G (eds) Ardipithecus kadabba. Late Miocene Evidence from the Middle Awash, Ethiopia. University of California Press, Berkeley, pp 469-516

Saegusa H, Hlusko LJ (2007) New late Miocene elephantoid (Mammalia: Proboscidea) fossils from Lemudong’o, Kenya. Kirtlandia 56:140-147

Sanders WJ (1997) Fossil Proboscidea from the Wembere-Manonga Formation, Manonga Valley, Tanzania. In: Harrison T (ed) Neogene Paleontology of the Manonga Valley, Tanzania. Plenum Press, New York, pp 265-310

Sanders WJ (2006) Comparative description and taxonomy of proboscidean fossils from Langebaanweg, South Africa. Afr Nat Hist 2:196-197

Sanders WJ (2007) Taxonomic review of fossil Proboscidea (Mammalia) from Langebaanweg, South Africa. Trans R Soc S Afr 62:1-16

Sanders WJ (2008) Review of fossil Proboscidea from the late Miocene-early Pliocene site of As Sahabi, Libya. Garyounis Sci Bull Special Issue No. 5:245-260

Sanders WJ (2011) Proboscidea. In: Harrison T (ed) Paleontology and Geology of Laetoli, Tanzania: Human Evolution in Context. Volume 2: Fossil Hominins and the Associated Fauna. Springer, Dordrecht, pp 233-262

Sanders WJ, Gheerbrant E, Harris J, Saegusa H, Delmer C (2010) Proboscidea. In: Werdelin L, Sanders WJ (eds) Cenozoic Mammals of Africa. University of California Press, Berkeley, pp 161-251

Semaw S, Simpson SW, Quade J, Renne PR, Butler RF, McIntosh WC, Levin N, Dominguez-Rodrigo M, Rogers MJ (2005) Early Pliocene hominids from Gona, Ethiopia. Nature 433:301-304

Tassy P (1986) Nouveaux Elephantoidea (Mammalia) dans le Miocène du Kenya. Cahiers de Paleontologie, Éditions du Centre de la Recherche Scientifique, Paris

Tassy P (1995) Les Proboscidiens (Mammalia) Fossiles du Rift Occidental, Ouganda. In: Pickford M, Senut B (eds) Geology and Palaeobiology of the Albertine Rift Valley, Uganda-Zaire. Vol. II: Palaeobiology. CIFEG Occasional Publications 1994/29, Orléans, pp 217-257

Tassy P (2003) Elephantoidea from Lothagam. In: Leakey MG, Harris JM (eds) Lothagam: The Dawn of Humanity in Eastern Africa. Columbia University Press, New York, pp 331-358

White TD, Moore RV, Suwa G (1984) Hadar biostratigraphy and hominid evolution. J Vertebr Paleontol 4:575-583