SUPPLEMENTARY MATERIAL
Key herbivores reveal limited functional redundancy on inshore coral reefs
C. L. Johansson 1, 2*, I. A. van de Leemput 3, M. Depczynski 4, A. S. Hoey 2
andD. R. Bellwood 2
Fig. S1. Mean abundance (100 m2) ± SE plotted against weighted mean reserve capacity ± SD for grazers on the Great Barrier Reef (GBR) (back, flat, crest and slope for inner, mid and outer reefs). The symbols on the GBR panels represent habitats, where ■ = back, ▲ = flat, ♦ = crest, ● = slope and ▼ = crest/slope. The colours of the symbols illustrate the different shelf locations. Grey zones represent the area which is less than the average abundance and 50 % reserve capacity (hence, one species is dominant) for each functional group. Habitats inside grey areas have less average abundance than the average for that functional group and are dominated by one species.
Table S1. Species included in the analysis. Locations where each species was recorded is highlighted in brackets, where N = Ningaloo Reef and G = Great Barrier Reef.
Browsers / Excavators / Grazers/Detritivores / ScrapersNaso annulatus [N, G] / Bolbometopon muricatum [G] / Acanthurus blochii [G] / Hipposcarus longiceps [N, G]
Naso brachycentron [G] / Cetoscarus bicolor [G] / Acanthurus dussumieri [N, G] / Scarus altipinnis [G]
Naso brevirostris [N, G] / Chlorurus bleekeri [G] / Acanthurus grammoptilus [N] / Scarus chamaeleon [N, G]
Naso lituratus [N, G] / Chlorurus microrhinos [N, G] / Acanthurus lineatus [G] / Scarus dimidiatus [N, G]
Naso tonganus [N, G] / Chlorurus sordidus [N, G] / Acanthurus nigricans [N] / Scarus festivus [G]
Naso unicornis [N, G] / Acanthurus nigricauda [N, G] / Scarus flavipectoralis [G]
Naso vlamingii [G] / Acanthurus nigrofuscus [N, G] / Scarus forsteni [G]
Platax orbicularis [N] / Acanthurus olivaceus [N] / Scarus frenatus [N, G]
Platax pinnatus [N] / Acanthurus triostegus [N, G] / Scarus ghobban [N, G]
Platax teira [N, G] / Acanthurus sp. [N] / Scarus globiceps [N, G]
Kyphosus cinerescens [G] / Acanthurus xanthopterus [G] / Scarus niger[G]
Kyphosus cornelii [N] / Zebrasoma scopas [N, G] / Scarus oviceps [N, G]
Kyphosus sydneyanus [N] / Zebrasoma veliferum [N, G] / Scarus prasiognathus [N]
Kyphosus vaigiensis [G] / Centropyge bicolor [G] / Scarus psittacus [N, G]
Leptoscarus vaigiensis [N] / Centropyge bispinosus [G] / Scarus rivulatus [N, G]
Siganus fuscesens [N] / Centropyge eibli [N] / Scarus rubroviolaceus [N, G]
Centropyge tibicen [N] / Scarus schlegeli [N, G]
Centropyge vrolicki [G] / Scarus sp. 1 [G]
Ctenochaetus binotatus [G] / Scarus sp. 2 [N]
Ctenochaetus striatus [N, G] / Scarus spinus [G]
Ctenochaetus strigosus [N]
Siganus argenteus [G]
Siganus corallinus [G]
Siganus doliatus [G]
Siganus laqueus [N]
Siganus lineatus [G]
Siganus puellus [G]
Siganus punctatissimus [G]
Siganus punctatus [G]
Siganus spinus [G]
Siganus sp. [N]
Siganus trispilos [N]
Siganus virgatus [N]
Siganus vulpinus [G]
Table S2. Results of Poisson logistic regressions comparing (a) species richness and (b) generic richness among three cross-shelf habitats on Ningaloo Reef. Regression coefficients and results of Tukey’s pairwise comparisons are given for each of the four functional groups. Significant results (p < 0.05) are given in italics.
Ningaloo Reef / Coefficients / Tukey's Post hocLagoon / Back / Slope / Lagoon * Back / Lagoon * Slope / Back * Slope
(a) Species Richness / Browsers / 0.8 / 2.6 / 2 / 0.004 / 0.028 / 0.378
Excavators / 0.3 / 1.2 / 0.7 / 0.032 / 0.220 / 0.257
Grazers / 1.3 / 3.9 / 4.3 / 0.001 / 0.000 / 0.659
Scrapers / 1.4 / 6.2 / 5.2 / 0.000 / 0.000 / 0.350
(b) Generic Richness / Browsers / 0.4 / 1.3 / 1.3 / 0.039 / 0.039 / 1.000
Excavators / 0.2 / 1 / 0.5 / 0.038 / 0.273 / 0.206
Grazers / 1 / 2.6 / 2.5 / 0.010 / 0.014 / 0.889
Scrapers / 0.6 / 1.3 / 1 / 0.117 / 0.323 / 0.533
Table S3. Results of Poisson logistic regressions comparing (a) species richness and (b) generic richness among eleven cross-shelf habitats on the GBR. Regression coefficients are given for each of the four functional groups.
CoefficientsInner / Mid / Outer
GBR / Back / Flat / S/C / Back / Crest / Flat / Slope / Back / Crest / Flat / Slope
(a) Species Richness / Browsers / 0.25 / 0.00 / 0.63 / 1.25 / 2.50 / 1.88 / 0.75 / 1.88 / 3.50 / 2.75 / 2.63
Excavators / 1.13 / 0.50 / 1.00 / 1.50 / 2.38 / 1.88 / 2.13 / 2.88 / 2.50 / 1.75 / 2.13
Grazers / 2.00 / 0.75 / 2.13 / 4.50 / 7.50 / 3.25 / 5.63 / 6.38 / 6.13 / 4.63 / 8.13
Scrapers / 1.88 / 2.25 / 2.00 / 5.88 / 6.88 / 6.25 / 6.88 / 8.50 / 5.38 / 4.88 / 6.75
(b) Generic Richness / Browsers / 0.25 / 0.00 / 0.63 / 0.75 / 1.25 / 0.88 / 0.75 / 1.00 / 1.13 / 1.00 / 1.25
Excavators / 0.88 / 0.50 / 1.00 / 1.13 / 1.25 / 1.13 / 1.38 / 1.88 / 1.75 / 1.38 / 1.13
Grazers / 1.63 / 0.75 / 1.75 / 2.00 / 4.13 / 1.88 / 3.50 / 3.75 / 4.13 / 2.63 / 4.75
Scrapers / 1.00 / 1.00 / 1.00 / 1.50 / 1.25 / 1.13 / 1.13 / 1.75 / 1.75 / 1.25 / 1.38
Table S4. P-values for species richness for (a) browsers, (b) excavators, (c) grazers and (d) scrapers the Great Barrier Reef (GBR). Results from Poisson Logistic Regression.Significant results (p < 0.05) are given in italics.
(a) / Inner / Mid / OuterBrowsers / Back / Flat / S/C / Back / Crest / Flat / Slope / Back / Crest / Flat / Slope
Inner / Back
Flat / 0.996
Slope/Crest / 0.273 / 0.995
Mid / Back / 0.038 / 0.995 / 0.206
Crest / 0.002 / 0.995 / 0.006 / 0.074
Flat / 0.007 / 0.995 / 0.033 / 0.321 / 0.400
Slope / 0.178 / 0.995 / 0.763 / 0.323 / 0.010 / 0.058
Outer / Back / 0.007 / 0.995 / 0.033 / 0.321 / 0.400 / 1.000 / 0.058
Crest / 0.000 / 0.995 / 0.000 / 0.005 / 0.250 / 0.051 / 0.001 / 0.051
Flat / 0.001 / 0.995 / 0.003 / 0.039 / 0.758 / 0.253 / 0.005 / 0.253 / 0.397
Slope / 0.001 / 0.995 / 0.004 / 0.053 / 0.876 / 0.320 / 0.007 / 0.320 / 0.319 / 0.879
(b) / Inner / Mid / Outer
Excavators / Back / Flat / S/C / Back / Crest / Flat / Slope / Back / Crest / Flat / Slope
Inner / Back
Flat / 0.177
Slope/Crest / 0.808 / 0.258
Mid / Back / 0.514 / 0.057 / 0.374
Crest / 0.065 / 0.005 / 0.040 / 0.213
Flat / 0.226 / 0.019 / 0.151 / 0.565 / 0.494
Slope / 0.123 / 0.009 / 0.079 / 0.356 / 0.739 / 0.724
Outer / Back / 0.017 / 0.001 / 0.010 / 0.068 / 0.538 / 0.198 / 0.345
Crest / 0.047 / 0.003 / 0.028 / 0.162 / 0.873 / 0.400 / 0.622 / 0.648
Flat / 0.301 / 0.027 / 0.207 / 0.695 / 0.386 / 0.853 / 0.591 / 0.143 / 0.306
Slope / 0.123 / 0.009 / 0.079 / 0.356 / 0.739 / 0.724 / 1.000 / 0.345 / 0.622 / 0.591
(c) / Inner / Mid / Outer
Grazers / Back / Flat / S/C / Back / Crest / Flat / Slope / Back / Crest / Flat / Slope
Inner / Back
Flat / 0.040
Slope/Crest / 0.862 / 0.028
Mid / Back / 0.007 / 0.000 / 0.011
Crest / 0.000 / 0.000 / 0.000 / 0.015
Flat / 0.127 / 0.001 / 0.173 / 0.206 / 0.000
Slope / 0.000 / 0.000 / 0.001 / 0.318 / 0.145 / 0.026
Outer / Back / 0.000 / 0.000 / 0.000 / 0.110 / 0.393 / 0.005 / 0.541
Crest / 0.000 / 0.000 / 0.000 / 0.160 / 0.293 / 0.009 / 0.680 / 0.841
Flat / 0.005 / 0.000 / 0.008 / 0.907 / 0.021 / 0.168 / 0.378 / 0.137 / 0.197
Slope / 0.000 / 0.000 / 0.000 / 0.004 / 0.655 / 0.000 / 0.058 / 0.195 / 0.135 / 0.006
(d) / Inner / Mid / Outer
Scrapers / Back / Flat / S/C / Back / Crest / Flat / Slope / Back / Crest / Flat / Slope
Inner / Back
Flat / 0.602
Slope/Crest / 0.857 / 0.732
Mid / Back / 0.000 / 0.001 / 0.000
Crest / 0.000 / 0.000 / 0.000 / 0.429
Flat / 0.000 / 0.000 / 0.000 / 0.761 / 0.626
Slope / 0.000 / 0.000 / 0.000 / 0.429 / 1.000 / 0.626
Outer / Back / 0.000 / 0.000 / 0.000 / 0.052 / 0.242 / 0.099 / 0.242
Crest / 0.000 / 0.002 / 0.001 / 0.673 / 0.227 / 0.468 / 0.227 / 0.019
Flat / 0.002 / 0.007 / 0.003 / 0.389 / 0.101 / 0.245 / 0.101 / 0.006 / 0.659
Slope / 0.000 / 0.000 / 0.000 / 0.486 / 0.924 / 0.695 / 0.924 / 0.206 / 0.265 / 0.121
Table S5. P-values for generic richness for (a) browsers, (b) excavators, (c) grazers and (d) scrapers the Great Barrier Reef (GBR). Results from Poisson Logistic Regression.Significant results (p < 0.05) are given in italics.
(a) / Inner / Mid / OuterBrowsers / Back / Flat / S/C / Back / Crest / Flat / Slope / Back / Crest / Flat / Slope
Inner / Back
Flat / 0.997
Slope/Crest / 0.273 / 0.997
Mid / Back / 0.178 / 0.997 / 0.763
Crest / 0.038 / 0.997 / 0.206 / 0.323
Flat / 0.118 / 0.997 / 0.566 / 0.782 / 0.469
Slope / 0.178 / 0.997 / 0.763 / 1.000 / 0.323 / 0.782
Outer / Back / 0.080 / 0.997 / 0.410 / 0.594 / 0.638 / 0.796 / 0.594
Crest / 0.054 / 0.997 / 0.292 / 0.442 / 0.819 / 0.618 / 0.442 / 0.808
Flat / 0.080 / 0.997 / 0.410 / 0.594 / 0.638 / 0.796 / 0.594 / 1.000 / 0.808
Slope / 0.038 / 0.997 / 0.206 / 0.323 / 1.000 / 0.469 / 0.323 / 0.638 / 0.819 / 0.638
(b) / Inner / Mid / Outer
Excavators / Back / Flat / S/C / Back / Crest / Flat / Slope / Back / Crest / Flat / Slope
Inner / Back
Flat / 0.372
Slope/Crest / 0.796 / 0.258
Mid / Back / 0.618 / 0.177 / 0.808
Crest / 0.469 / 0.121 / 0.638 / 0.819
Flat / 0.618 / 0.177 / 0.808 / 1.000 / 0.819
Slope / 0.350 / 0.083 / 0.493 / 0.655 / 0.827 / 0.655
Outer / Back / 0.096 / 0.019 / 0.151 / 0.226 / 0.321 / 0.226 / 0.435
Crest / 0.134 / 0.027 / 0.207 / 0.301 / 0.416 / 0.301 / 0.549 / 0.853
Flat / 0.350 / 0.083 / 0.493 / 0.655 / 0.827 / 0.655 / 1.000 / 0.435 / 0.549
Slope / 0.618 / 0.177 / 0.808 / 1.000 / 0.819 / 1.000 / 0.655 / 0.226 / 0.301 / 0.655
(c) / Inner / Mid / Outer
Grazers / Back / Flat / S/C / Back / Crest / Flat / Slope / Back / Crest / Flat / Slope
Inner / Back
Flat / 0.117
Slope/Crest / 0.847 / 0.082
Mid / Back / 0.578 / 0.040 / 0.715
Crest / 0.004 / 0.000 / 0.007 / 0.017
Flat / 0.706 / 0.058 / 0.853 / 0.857 / 0.011
Slope / 0.022 / 0.001 / 0.034 / 0.074 / 0.523 / 0.051
Outer / Back / 0.012 / 0.000 / 0.019 / 0.042 / 0.706 / 0.028 / 0.793
Crest / 0.004 / 0.000 / 0.007 / 0.017 / 1.000 / 0.011 / 0.523 / 0.706
Flat / 0.174 / 0.007 / 0.240 / 0.413 / 0.105 / 0.320 / 0.319 / 0.210 / 0.105
Slope / 0.001 / 0.000 / 0.001 / 0.004 / 0.553 / 0.002 / 0.220 / 0.333 / 0.553 / 0.029
(d) / Inner / Mid / Outer
Scrapers / Back / Flat / S/C / Back / Crest / Flat / Slope / Back / Crest / Flat / Slope
Inner / Back
Flat / 1.000
Slope/Crest / 1.000 / 1.000
Mid / Back / 0.374 / 0.374 / 0.374
Crest / 0.638 / 0.638 / 0.638 / 0.670
Flat / 0.808 / 0.808 / 0.808 / 0.514 / 0.819
Slope / 0.808 / 0.808 / 0.808 / 0.514 / 0.819 / 1.000
Outer / Back / 0.207 / 0.207 / 0.207 / 0.695 / 0.416 / 0.301 / 0.301
Crest / 0.207 / 0.207 / 0.207 / 0.695 / 0.416 / 0.301 / 0.301 / 1.000
Flat / 0.638 / 0.638 / 0.638 / 0.670 / 1.000 / 0.819 / 0.819 / 0.416 / 0.416
Slope / 0.493 / 0.493 / 0.493 / 0.835 / 0.827 / 0.655 / 0.655 / 0.549 / 0.549 / 0.827
Table S6. Results for abundance, from one-way ANOVA and Tukey’s Post Hoc test, to test for significant differences between habitats on Ningaloo Reef, for each functional group.Significant results (p < 0.05) are given in italics.
Tukey's Post hocdf / SS / MS / F / p / Lagoon * Back / Lagoon * Slope / Back * Slope
Browsers
Habitat / 2 / 0.001 / 0.001 / 0.695 / 0.508 / 0.781 / 0.872 / 0.478
Error / 27 / 0.024 / 0.001
Total / 29 / 0.026
Excavators
Habitat / 2 / 0.001 / 0.001 / 0.328 / 0.724 / 0.879 / 0.946 / 0.704
Error / 27 / 0.043 / 0.002
Total / 29 / 0.044
Grazers
Habitat / 2 / 0.019 / 0.009 / 7.471 / 0.003 / 0.002 / 0.135 / 0.165
Error / 27 / 0.034 / 0.001
Total / 29 / 0.053
Scrapers
Habitat / 2 / 0.004 / 0.002 / 7.724 / 0.002 / 0.005 / 0.007 / 0.990
Error / 27 / 0.007 / 0.000
Total / 29 / 0.011
Table S7. Results for abundance, from one-way ANOVA, to test for significant differences between habitats on the Great Barrier Reef (GBR), for each functional group.Significant results (p < 0.05) are given in italics.
df / SS / MS / F / pBrowsers
Habitat / 10 / 0.025 / 0.002 / 6.576 / 0.000
Error / 77 / 0.029 / 0.000
Total / 87 / 0.054
Excavators
Habitat / 10 / 0.012 / 0.001 / 6.190 / 0.000
Error / 77 / 0.015 / 0.000
Total / 87 / 0.027
Grazers
Habitat / 10 / 5.464 / 0.546 / 10.936 / 0.000
Error / 77 / 3.847 / 0.050
Total / 87 / 9.311
Scrapers
Habitat / 10 / 0.047 / 0.005 / 3.541 / 0.001
Error / 77 / 0.103 / 0.001
Total / 87 / 0.150
Table S8. Significant differences in abundance, from Tukey’s Post Hoc test, between habitats on the Great Barrier Reef (GBR);for (a) browsers, (b) excavators, (c) grazers and (d) scrapers. Significant results (p < 0.05) are given in italics.
(a) / Inner / Mid / OuterBrowsers / Back / Flat / Crest/Slope / Back / Crest / Flat / Slope / Back / Crest / Flat / Slope
Inner / Back
Flat / 1.000
S/C / 1.000 / 1.000
Mid / Back / 1.000 / 1.000 / 1.000
Crest / 0.592 / 0.547 / 0.710 / 0.837
Flat / 0.895 / 0.868 / 0.949 / 0.985 / 1.000
Slope / 1.000 / 1.000 / 1.000 / 1.000 / 0.724 / 0.954
Outer / Back / 0.932 / 0.911 / 0.971 / 0.993 / 1.000 / 1.000 / 0.974
Crest / 0.002 / 0.002 / 0.004 / 0.009 / 0.502 / 0.201 / 0.005 / 0.159
Flat / 0.000 / 0.000 / 0.000 / 0.000 / 0.048 / 0.010 / 0.000 / 0.007 / 0.989
Slope / 0.847 / 0.814 / 0.918 / 0.971 / 1.000 / 1.000 / 0.925 / 1.000 / 0.251 / 0.014
(b) / Inner / Mid / Outer
Excavators / Back / Flat / Crest/Slope / Back / Crest / Flat / Slope / Back / Crest / Flat / Slope
Inner / Back
Flat / 1.000
S/C / 1.000 / 1.000
Mid / Back / 0.402 / 0.090 / 0.112
Crest / 0.000 / 0.000 / 0.000 / 0.207
Flat / 0.048 / 0.005 / 0.007 / 0.996 / 0.800
Slope / 0.669 / 0.220 / 0.263 / 1.000 / 0.083 / 0.953
Outer / Back / 0.383 / 0.083 / 0.104 / 1.000 / 0.220 / 0.997 / 1.000
Crest / 0.149 / 0.022 / 0.029 / 1.000 / 0.503 / 1.000 / 0.998 / 1.000
Flat / 0.953 / 0.586 / 0.649 / 0.995 / 0.015 / 0.669 / 1.000 / 0.994 / 0.910
Slope / 0.930 / 0.523 / 0.586 / 0.998 / 0.020 / 0.728 / 1.000 / 0.997 / 0.938 / 1.000
(c) / Inner / Mid / Outer
Grazers / Back / Flat / Crest/Slope / Back / Crest / Flat / Slope / Back / Crest / Flat / Slope
Inner / Back
Flat / 1.000
S/C / 1.000 / 1.000
Mid / Back / 1.000 / 1.000 / 1.000
Crest / 1.000 / 0.999 / 1.000 / 1.000
Flat / 0.999 / 0.997 / 0.999 / 1.000 / 1.000
Slope / 1.000 / 1.000 / 1.000 / 1.000 / 1.000 / 1.000
Outer / Back / 0.492 / 0.407 / 0.479 / 0.689 / 0.916 / 0.938 / 0.601
Crest / 0.000 / 0.000 / 0.000 / 0.000 / 0.000 / 0.000 / 0.000 / 0.003
Flat / 0.000 / 0.000 / 0.000 / 0.000 / 0.001 / 0.001 / 0.000 / 0.087 / 0.985
Slope / 0.034 / 0.023 / 0.032 / 0.074 / 0.206 / 0.240 / 0.052 / 0.977 / 0.104 / 0.730
(d) / Inner / Mid / Outer
Scrapers / Back / Flat / Crest/Slope / Back / Crest / Flat / Slope / Back / Crest / Flat / Slope
Inner / Back
Flat / 1.000
S/C / 1.000 / 1.000
Mid / Back / 1.000 / 1.000 / 1.000
Crest / 1.000 / 1.000 / 1.000 / 1.000
Flat / 0.818 / 0.929 / 0.830 / 0.958 / 0.961
Slope / 1.000 / 1.000 / 1.000 / 1.000 / 1.000 / 0.933
Outer / Back / 0.002 / 0.005 / 0.002 / 0.008 / 0.008 / 0.262 / 0.005
Crest / 1.000 / 1.000 / 1.000 / 1.000 / 1.000 / 0.993 / 1.000 / 0.019
Flat / 1.000 / 0.999 / 1.000 / 0.996 / 0.996 / 0.432 / 0.998 / 0.000 / 0.971
Slope / 1.000 / 1.000 / 1.000 / 1.000 / 1.000 / 0.906 / 1.000 / 0.004 / 1.000 / 0.999
Table S9. Variations in weighted reserve capacity among habitats on Ningaloo Reef.Coefficients and P-values from weighted binomial Logistic Regression with abundance (the sample size from which reserve capacity is calculated) as the weight.Significant results (p < 0.05) are given in italics.
Ningaloo Reef / Coefficients / Tukey's Post hocLagoon / Back / Slope / Lagoon * Back / Lagoon * Slope / Back * Slope
Reserve Capacity / Browsers / 0.11 / 0.26 / 0.23 / 0.000 / 0.004 / 0.460
Excavators / 0.03 / 0.01 / 0.02 / 0.091 / 0.401 / 0.515
Grazers / 0.27 / 0.18 / 0.49 / 0.047 / 0.000 / 0.000
Scrapers / 0.34 / 0.60 / 0.55 / 0.000 / 0.005 / 0.152
Table S10.Variations in weighted reserve capacity among habitats on the Great Barrier Reef (GBR).Coefficients from weighted binomial Logistic Regression with abundance (the sample size from which reserve capacity is calculated) as the weight.
CoefficientsInner / Mid / Outer
GBR / Back / Flat / S/C / Back / Crest / Flat / Slope / Back / Crest / Flat / Slope
Reserve Capacity / Browsers / 0.33 / - / 0.27 / 0.19 / 0.36 / 0.32 / 0.00 / 0.32 / 0.55 / 0.44 / 0.25
Excavators / 0.06 / 0.00 / 0.00 / 0.07 / 0.23 / 0.19 / 0.15 / 0.30 / 0.25 / 0.13 / 0.29
Grazers / 0.36 / 0.15 / 0.30 / 0.58 / 0.57 / 0.31 / 0.59 / 0.48 / 0.29 / 0.29 / 0.69
Scrapers / 0.09 / 0.15 / 0.09 / 0.63 / 0.58 / 0.64 / 0.64 / 0.47 / 0.54 / 0.53 / 0.55
Table S11. P-values for variations in weighted reserve capacity among habitats on the Great Barrier Reef (GBR); for (a) browsers, (b) excavators, (c) grazers and (d) scrapers. Results from weighted binomial Logistic Regression with abundance (the sample size from which reserve capacity is calculated) as the weight.Significant results (p < 0.05) are given in italics.
(a) / Inner / Mid / OuterBrowsers / Back / Flat / Crest/Slope / Back / Crest / Flat / Slope / Back / Crest / Flat / Slope
Inner / Back
Flat / 0.837
S/C / 0.837 / -
Mid / Back / 0.575 / - / 0.595
Crest / 0.932 / - / 0.579 / 0.148
Flat / 0.974 / - / 0.732 / 0.241 / 0.654
Slope / 0.990 / - / 0.990 / 0.990 / 0.990 / 0.990
Outer / Back / 0.958 / - / 0.760 / 0.262 / 0.607 / 0.944 / 0.990
Crest / 0.473 / - / 0.090 / 0.004 / 0.002 / 0.001 / 0.989 / 0.001
Flat / 0.704 / - / 0.273 / 0.032 / 0.141 / 0.068 / 0.990 / 0.064 / 0.029
Slope / 0.756 / - / 0.889 / 0.551 / 0.139 / 0.332 / 0.990 / 0.377 / 0.000 / 0.003
(b) / Inner / Mid / Outer
Excavators / Back / Flat / Crest/Slope / Back / Crest / Flat / Slope / Back / Crest / Flat / Slope
Inner / Back
Flat / 0.995
S/C / 0.994 / 1.000
Mid / Back / 0.779 / 0.995 / 0.994
Crest / 0.032 / 0.994 / 0.993 / 0.001
Flat / 0.070 / 0.995 / 0.993 / 0.007 / 0.391
Slope / 0.171 / 0.995 / 0.993 / 0.069 / 0.101 / 0.384
Outer / Back / 0.010 / 0.994 / 0.993 / 0.000 / 0.192 / 0.050 / 0.011
Crest / 0.022 / 0.994 / 0.993 / 0.000 / 0.642 / 0.227 / 0.057 / 0.439
Flat / 0.286 / 0.995 / 0.993 / 0.208 / 0.048 / 0.195 / 0.630 / 0.006 / 0.027
Slope / 0.013 / 0.994 / 0.993 / 0.000 / 0.307 / 0.098 / 0.024 / 0.890 / 0.570 / 0.012
(c) / Inner / Mid / Outer
Grazers / Back / Flat / Crest/Slope / Back / Crest / Flat / Slope / Back / Crest / Flat / Slope
Inner / Back
Flat / 0.086
S/C / 0.434 / 0.191
Mid / Back / 0.000 / 0.001 / 0.000
Crest / 0.000 / 0.002 / 0.000 / 0.708
Flat / 0.410 / 0.137 / 0.817 / 0.000 / 0.000
Slope / 0.000 / 0.001 / 0.000 / 0.854 / 0.591 / 0.000
Outer / Back / 0.025 / 0.008 / 0.002 / 0.005 / 0.002 / 0.000 / 0.008
Crest / 0.171 / 0.185 / 0.847 / 0.000 / 0.000 / 0.283 / 0.000 / 0.000
Flat / 0.196 / 0.176 / 0.898 / 0.000 / 0.000 / 0.367 / 0.000 / 0.000 / 0.767
Slope / 0.000 / 0.000 / 0.000 / 0.001 / 0.000 / 0.000 / 0.007 / 0.000 / 0.000 / 0.000
(d) / Inner / Mid / Outer
Scrapers / Back / Flat / Crest/Slope / Back / Crest / Flat / Slope / Back / Crest / Flat / Slope
Inner / Back
Flat / 0.075
S/C / 0.839 / 0.048
Mid / Back / 0.000 / 0.000 / 0.000
Crest / 0.000 / 0.000 / 0.000 / 0.324
Flat / 0.000 / 0.000 / 0.000 / 0.677 / 0.134
Slope / 0.000 / 0.000 / 0.000 / 0.798 / 0.218 / 0.895
Outer / Back / 0.000 / 0.000 / 0.000 / 0.000 / 0.004 / 0.000 / 0.000
Crest / 0.000 / 0.000 / 0.000 / 0.043 / 0.307 / 0.008 / 0.024 / 0.079
Flat / 0.000 / 0.000 / 0.000 / 0.053 / 0.289 / 0.015 / 0.032 / 0.211 / 0.859
Slope / 0.000 / 0.000 / 0.000 / 0.098 / 0.492 / 0.027 / 0.059 / 0.045 / 0.764 / 0.665