1

Appendix 1 Voucher details for the specimens examined. Institutional abbreviations: AMNH, American Museum of Natural History, New York, New York, USA;MCZ,Museum of Comparative Zoology, Harvard University, Cambridge, Massachusetts, USA; BM, The Natural History Museum, London, UK; ZMUC, Natural History Museum of Denmark (Zoological Museum), Copenhagen, Denmark.

Akymnopellis chilensis (Gervais, 1847)—Argentina: TucumánProvince: Horco Molle, Cerro San Javier; 16.ix.2001; C. Mattoni; AMNH, MCZ DNA100473

Alipes crotalus(Gerstaecker, 1845);Central African Republic; xii.1998; J. Wenzel; MCZ DNA100273

Alipes grandidieri Lucas, 1864—Tanzania; purchased from commercial pet trade, 2010

Arthrorhabdus formosus Pocock 1891—33º35'9.9"S 22º10'52.9E; South Africa: Western Cape Province: Oudtshoom District,western boundary of Oudtshoom town; 3.xii.2006; I. Engelbrecht, R. Mercurio; AMNHLP 6656, MCZ DNA104693.South Africa: Klipfontein, Namaqualand; BM 1904.9.1.29-31

Asanada brevicornis Meinert, 1886—Thailand: NakhonSawanProvince, Nongbua district, Tam Pra; 8.x.2009; H. Enghoff; ZMUC, MCZ DNA 104867. Andamans; E.W. Oates: BM 1889.7.15.21-23

Asanada sokotrana Pocock, 1899—12º35'53.8''N 14º9'59.4''W;Guinea-Bissau: 3 km S of Pirada at the border,secondary broadleaf forest; 28.vi.2005; J. Huff, V. Vignoli; AMNH LP 4878; MCZ DNA104721

Campylostigmus decipiens Ribaut, 1923—22º11'S 166º01'E; New Caledonia: Mt Koghis; 28.xi.2001; G.B. Monteith

Cormocephalus aurantiipes(Newport, 1844)—32º45'45''S 116º04'36''E; Australia: Western Australia: Lane Poole Reserve, S. of Dwellingup; 21.i.2006; G.D. Edgecombe, G. Giribet; MCZ DNA103951

Cormocephalus gervaisianusC.L. Koch—36º42'530''N 10º20'680''E; Tunisia: Tunis District, Bou Kournine N.P.Thuya, Eucalyptus/ dry river bed, under stones and logs; 4.iii.2008; P. Stoev, N. Akkari; MCZ DNA104642

Cryptops doriaePocock, 1891—18º35'7''N 98º28'59''E;Thailand: ChiangMaiProvince: Chomthong district, Doi Inthanon near summit; 12.x.2009; H. Enghoff; ZMUC, MCZ DNA104871

Cryptops punicusSilvestri, 1896—40º25'36''N 8º59'57''E; Italy: Sardinia: Monte Rasu, near Bono, Sassari; 25.iii.2008; G.D. Edgecombe, G. Giribet; MCZ DNA104681

Cryptops trisulcatusBrolemann, 1902—37º54'17''N 15º04'37''E; Italy: Sicily: road from Francavilla di Sicilia to Randazzo, Catania; 20.iii.2008; G.D. Edgecombe, G. Giribet, Z. Johanson; MCZ DNA104689

Dinocryptops miersiiNewport, 1845—Brazil: ParanáState:Usina Santa Clara; 21.xii.2003; Carlos Edwardo Conte; MCZ DNA101160

Ethmostigmus rubripes(Brandt, 1840)—Australia: New South Wales:Merriganowry, Cowra Shire; viii.1995; G.D. Edgecombe; MCZ DNA100277

Ethmostigmus trigonopodus(Leach, 1817)—Approx. 02°28.929 S, 016°30.371 E; Democratic Republic of Congo:BandunguProvince: Malebo field station; x.2010; MCZ DNA106002

Hemiscolopendra marginataSay, 1821—37°14'31.91''N 78°27'52.02''W; USA: Virginia: Hampden-SydneyCollege campus, PrinceEdwardCounty; 24.ix.2009; W.A. Shear; MCZ DNA104671

Otostigmus astenus(Kohlrausch, 1878)—17°45'18.2S 167°20'19.2'' E; Vanuatu: EfateIsland: private conservation area, road to Erakor;16.viii.2008; P. Sharma, J. Murienne; MCZ DNA103943

Otostigmus rugulosus Porat, 1876—Burma: Rangoon; E.W. Oates; BM 89.7.15.34.

Otostigmus spinicauda(Newport, 1844)—Spain: Canary Islands:Banco Cervo, Fuerteventura; 2.xii.2001; P. Oromí; MCZ DNA104614

Rhysida nuda(Newport, 1845)—Australia: New South Wales:ca 28 km S of Nundle; 6.vii.1998; W. Ponder; AM KS 58456, MCZ DNA100278

Scolopendra alternans Leach, 1813—24º42'353''N 81º22'822''W; USA: Florida: Monroe Co., Big Pine Key, off Key Deer Blvd, Blue Hole; 17.xi.2009; G. Giribet, P. Sierwald; MCZ DNA104817

Scolopendra canidens Newport, 1844—32°55'506''N 10°26'913''E;Tunisia: Tataouine District, surroundings of Tataouine city; arid biotope, slope, stones, scattered trees of Pinus (planted), under stones;13.iii.2008; P. Stoev, N. Akkari; MCZ DNA104640

Scolopendra laeta Haase, 1887—Australia: New South Wales: Cowra Shire, Mooroobin property, 1.5 km NW Merriganowry, GR 642750 6373750, under rocks in paddock;24.ix.2005; G.D. Edgecombe; MCZ DNA103954

Scolopendra morsitans Linnaeus, 1758—26º54'44''S 31º53'11''E; Swaziland: Big Bend, Mlawula Nature Reserve; 3.iv.2001; G. Giribet, L. Prendini; MCZ DNA100677

Scolopendra oraniensis Lucas, 1846—39°18'14''N 9°22'57''E;Italy: Sardinia: route SS125, near S. Gregorio; 23.iii.2008; G.D.Edgecombe, G. Giribet; MCZ DNA104684

Scolopendra subspinipes subspinipes Leach, 1815—17°45'18.2''S 167°20'19.2''E;Vanuatu: EfateIsland: private conservation area, road to Erakor; 16.viii.2008; P. Sharma, J. Murienne; MCZ DNA103942

Scolopendra valida Lucas, 1840—Spain: San Andrés, Tenerife; 4.iv.2008;Spain; P. Oromí; MCZ DNA104872

Scolopocryptops melanostomaNewport, 1845—18°4'15''S 178°26'39.9''E; Fiji:Viti LevuIsland: Savura park along ridge trail; 31.vii.2008; J. Murienne, P. Sharma; MCZ DNA104006

Scolopocryptops mexicanus Humbert & Saussure, 1869—Mexico: Omilteme, Guerrero; Godman, Salvin; BM 97.3.1.85-95+

Scolopocryptops rubiginosus L. Koch, 1878—24º45'44''N 121º35'57''E; Taiwan: Llan County, Fushan Botanical Garden, primary forest; 22.v.2010; Z. Korsós, Y. Nakamura; MCZ DNA105912

Theatops erythrocephalus(C.L. Koch, 1847)—Spain: Barcelona Province; xi.1995; A. Serra; MCZ DNA103996

Theatops posticus (Say, 1821)—USA: Texas: Four Corner Trail, Radcliff Recreation Area, 18 NE Crockett; 3.i.2003; C.T. McAllister, G. Torres; BM

Zoomorphology

Spiracle structure in scolopendromorph centipedes (Chilopoda: Scolopendromorpha) and its contribution to phylogenetics

V. Vahtera

Museum of Comparative Zoology, Department of Organismic and Evolutionary Biology, HarvardUniversity,

26 Oxford Street, Cambridge, MA02138, USA

Current address: Finnish Museum of Natural History, Zoology Unit,

P.O. Box 17, 00014 University of Helsinki, Helsinki, Finland

e-mail:

Tel.: +358919128857

G. D. Edgecombe

Department of Palaeontology,

The NaturalHistoryMuseum,

Cromwell Road, LondonSW7 5BD, UK

G. Giribet

Museum of Comparative Zoology, Department of Organismic and Evolutionary Biology, HarvardUniversity,

26 Oxford Street, Cambridge, MA02138, USA