Subkingdom Parazoa

Phylum Porifera

Habitat

5000 species (All aquatic, some brackish, most marine)

Anatomy

Sessile
Two layers imbedded in a gelatinous matrix (mesohyl)
Outer layer of epidermal cells
Inner layer of choanocytes or epidermal cells
Amoebocytes and spicules in mesohyl
Skeleton
Spicules of Calcium carbonate or silicon
Spongin is a flexible skeleton
Cells
Porocytes -- Form incurrent pores in the epithelial body wall
Choanocytes
Flagellated cell
Contain collars
Create water current and trap food
Amoebocytes
Wandering cells in the mesohyl
Can differentiate into any other cell type
Functions

Food digestion

Nutrient distribution

Food storage

Gamete formation

Spicule formation

Epidermal cells
Line the outside and sometimes the spongocoel and canals
Basic body plan
Choanocytes on inside create a water current

Water enters incurrent pores

Pores are formed by porocytes

Goes into the spongocoel

Central cavity of the sponge

Exits the sponge through oscula

Large excurrent opening of the spongocoel

Filter feeders

All cells capable of phagocytosis / intracellular digestion and absorption.
Feeding

Choanocytes

Collar sieve with mucous

Creates water current and traps food

Slides down and phagocytized at base

Amoebocytes

Collects, digests, distributes, and stores food

Other physiology

No respiratory or excretory organs
Gas exchange and ammonia removed by water flow

Reproduction

Asexual

Regeneration powers

Budding (rare)

Gemmules

Gemmules remain dormant during droughts and freezes

Released during good conditions

Sexual

General

Monoecious more common than dieocious

Viviparous more common that oviparous

Gametes

Amoebocytes and choanocytes can differentiate into egg and sperm

Egg remains in mesohyl

Eggs may engulf nurse cells to accumulate nutrients

Sperm is released

Fertilization

Sperm is phagocytized by choanocytes of another sponge

Choanocyte delivers sperm to egg

Choanocytes releases sperm nuclei or both are engulfed

Larvae

Zygote develops into a flagellated larvae which escapes through the osculum

Most have parenchymella larvae

Cilia all over

Some have amphiblastula

cilia over one end

Metamorphosis

Inversion similar in gastrulation

Flagella now located on the inside

Subkingdom Eumetazoa

Branch Radiata

Characteristics

Radial symmetry
Diploblastic

Phylum Cnidaria

Gastrovascular cavity with 1 opening

Polyp (sessile-attached by aboral end) and medusa (floating - mouth/anus facding down) body forms

Tentacles with cnidocytes with nematocysts (stingers) to capture prey and for defense

Carnivorous

Class Scyphozoa --jellyfish; class Anthozoa -- sea anemones and corals (secrete shell); class Hydrozoa

Branch Bilateria

Acoelomates

Characteristics

Triploblastic (ectoderm, endoderm, and mesoderm)

Bilateral symmetry

Acoelomates

No fluid-filled body cavity

Phylum Platyhelminthes (Flatworms)

Poss several organs

Class Turbellaria

Free living

Carnivorous

Rudimentary brain

Class Trematoda (flukes)

All parasites with two or more hosts in their life cycle

Class Cestoda (tapworms)

All parasites of vertebrate’s digestive system

No digestive system

Pseudocoelomates

Fluid-filled body cavity not completely lined with mesoderm

Phylum Nematoda (roundworms)

Dioecious (separate sexes)

Important plant and animals pathogens

Coelomates

Characteristics

Contain a true coelom (body cavity completely lined with mesoderm)

Coelom used as a hydrostatic skeleton, for storage, and to cushoin organs

Protostomes

Characteristics

First evagination (blastopore) of gastrula forms mouth

Coelom forms by splitting of mesoderm

Spiral and determinant cleavage

Phylum Mollusca

Characteristics

Mostly marine

3 distinct body parts

Visceral mass - soft (contains organs)

Foot - muscular (movement)

Mantle - covers visceral mass (secretes shell)

Many show cephalization

Class Gastropoda

Slugs and snails

Herbivorous

Torsion

One side of visceral mass grows faster than other

Results in anus being above mouth

Use radula for feeding / rasping

Class Bivalvia

Clams (moves with foot), oysters, mussels, and scallops (moves by flapping shell)

2 shells hinged at the back, kept colesed by adductor muscle

Filter feeders

Class Cephalopoda

Octopuses and squid

Carnivorous (highly cephalized -- need to be to locate and capture prey)

Squid - reduced internal shell, open sea, jet propulsion

Octopus - no shell, ground dwelling, moves with tentacles

Phylum Annelidia

Characteristics

Segmented worms

Segments partitioned by septa - blood vessel, dorsal nerve and digestive tract transcend segments

Complete digestive tract

Cutaneous respiration

Hermaphroditic

Class oligochaeta

Earthworms

Important in tilling earth (1 acres may contain 50K worms, tilling 18 tons per year)

Class Polychaeta

Contain one pair of parapodia per segment

Parapodia function in locomotion and gas exchange

Probable gave rise to arthropods (parapodia became specialized appendages)

Class Hirudinea

Leeches (exoparasites)

Phylum Arthropoda

Characteristics

Most successful animal phylum

Species diversity (1 million species / 3/4 all animal species)

Distribution (almost everywhere)

Number of individuals (1 billion billion)

Probably evolved from the annelids

Number of segments became reduced

Segments became specialized (walking, feeding, sensory, copulation and defense)

Contain a hard exoskeleton (protein and chitin)

Protection

Muscle attachment

Water conservation - -important for movement onto land

Support -- important for movement onto land

Must molt (exoskeleton does not grow)

Extensive cephalization (eyes, olfactory receptors, antennae - for touch & smell)

Open circulatory system with hemolymph

Compete with humans for food and spread diseases

Essential for pollination of many flowers and produce many unique and useful products (honey and silk)

Efficient breathing apparatus (gills, trachea, book lungs)

Metamorphosis is common (reduces competition between juveniles and adults)

Subphylum Chelicerata

Characteristics

6 pairs of appendages including chelicerae (fang-like appendages used in feeding) and pedipalps

No attennae or mandibles

Mostly terrestrial

Class Arachnida

Scorpions, spiders, mites, and ticks

6 pairs of appendages

Chelicerae, pedipalps (sensing and feeding), 4 pairs of legs

Feedign (all predacious)

Fang-like chelicerae

Poison glands

Secrete digestive juices

Suck up liquid soup

Book lungs (unique to this class) and trachea system

Malpighian tubules for secretion (empties urine into digestive tract - nearly dry urine/feces mix)

Eight simple eyes

Silk

Protein made by abdominal glands

Spinnerets spin silk into fibers

Innate ability to produce webs, drop lines, wrap eggs, make gifts (courting rituals)

Scorpions

Feed on insects and spiders, can sense vibrations, has stinger on tails)

Ticks and Mites

Spread diseases like Rocky Mountain spotted fever and Lyme disease / chiggers

Subphylum Uniramia

Characteristics

Claw-like chelicerae replaced with jaw-like mandibles

Pedipalps replaced with maxillae for food handling

Have 1 pair of antennae

Class Diplopoda -- Millipedes

2 pairs of legs per segment

Herbivores

Earliest land animals (ate mosses)

Class Chilopoda -- Centipedes

1 pair of legs per segment

Carnivores (earthworms, cockroaches, and insects)

Insecta

Entomology - scientific study of insects

Most successful of all arthropod groups

Flight

1 or 2 pairs of wings (from cuticle, not appendages like bats and birds)

Escape, finding mates and food, dispersal

3 pairs of walking legs

Gas exchange by trachea (branching tubes) and spiracles (opening)

Metamorphosis

Complete (drastic change)

(egg  larvae  pupil  adult)caterpillar - cocoon - butterfly)

Incomplete

(egg  immature nymphs  adult)

Can be vectors for disease (malaria, African sleeping sickness, etc)

Subphylum Crustacea

Lobsters, crayfish (fresh water), crabs, shrimp, krill, barnacles

Sow bugs and pill bugs (rolly pollies) are terrestrial

Separate sexes

2 pairs of antennae

Deuterostomes

Characteristics

Radial indeterminate cleavage

Blastopore forms anus

Coelom forms from endoderm outpocketing

Phylum Echinodermata

Characteristics

No cephalization or segmentation

Calcareous endoskeleton covered with skin (skin protected with pincer like pedicellariae)

Separate sexes

Sessile or sedentary

Adults have radial symmetry

Juveniles metamorphosize from bilateral symmetry to radial symmetric adults

Water vascular system (locomotion, feeding and gas exchange) from coelom

Fluid moved by cilia

Used for gas exchange, waste removal, movement and feeding

All marine

Strong powers of regeneration

Class Asteroides -- sea stars (feed on mollusks)

Class Ophiuroidea -- brittle stars (flexible arms); scavengers

Class Crinoidea -- sea lilies (often attached filter feeders)

Class Holothuroidea --sea cucumbers (lost shell) -- deposit feeders in mucky areas

Class Echniodea -- sea urchins and sand dollars (feed on seaweed)

Phylum Chordata

Characteristics

Notochord (in all chordate embryos)

Longitudinal, flexible rod

Between gut and nerve chord

Serves as skeleton in some vertebrates

Reduced to pads between vertebrates in humans

Hollow-dorsal nerve chord

Ectoderm rolls in a tube dorsal to notochord

Other phyla have solid nerve chord, usually ventral

Forms central nervous system (brain and spinal cord)

Pharyngeal slits (in all chordate embryos)

Slits in pharynx

Filter feeding

Gas exchange

Muscular post-anal tail

Subphylum Cephalochordata

Subphylum Urochordata

Subphylum Vertebrata

Characteristics

Sexually mature larvae

Highly cephalized

Nerve chord protected with vertebrate

2 pairs of appendages (fins, wings, legs, etc.)

Living endoskeleton (contains osteocytes - skeleton can grow)

Closed circulatory system (blood stays in vessels)

Waste removed with kidneys

Dioecious - separate sexes (sexual reproduction)

Class Agnatha (jawless fish)

Uses mouth to latch onto fish, rasps hole in fish, and feeds on blood

Class Chondrichthyes (cartilaginous fish) - sharks and rays

Carnivorous (exceptions: whale shark - filter feeder)

Good senses (electrical impulses, water pressure, etc.)

Oviparous (young hatch from eggs outside body); ovoviviparous (young hatch from eggs in the mother’s uterus); or viviparous (young are nourished by placenta)

Class Osteichthyes (bony fish)

Swim bladder (fills with gas - from blood) to let fish float

Operculum (covers gills - brings water in mouth and out gills)

Oviparous

Most numerous of all vertebrates

Class Amphibia

Three orders

Salamanders(aquatic, land, or both)

Caecilians(worm-like -- tropics)

Toads and frogs(metamorphosis)

Iinefficient lungs; cutaneous respiration

Metamorphosis:young do not compete with adults for resources

Tadpoleherbivores (aquatic)

Adultsinsects (land or aquatic)

External fertilization

Class Reptilia

Keratin - water proof skin

obtain oxygen from lungs

Oviparousinternal fertilization

shell on amniote egg

Ectothermic“cold-blooded”

energy conservation (10% the food requirement of mammals)

Evolution (one branch contained the turtles, snakes and lizards)

(other branch contained the dinosaurs, crocodiles, and alligators - this branch gave rise to the mammals and latter the birds)

ascended from amphibians

gave rise to birds and mammals

3 extant orders:

lizards and snakes

turtles

alligators and crocodiles

Dinosaurs as quick moving endotherms (went extinct around time gymnosperms gave rise to angiosperms - mammals radiated when dinosaurs became extinct)

Class Aves

Evolved from reptiles

retained amniote egg and scales

Hollow bones, toothless, absence of some organs (example - only one ovary)

Endothermic

Oviparous

internal fertilization (though most birds have no penis)

eggs kept warm

Feathers from keratin (hair, fingernails, scales)

Ratite birds (flightless - lacking breast muscle - ostriches)

Carinate birds (flight birds - includes penguins)

Class Mammalia

-from reptiles (before birds)

Hair (insulation)

4 chambered heart (keeps oxygenated and deoxygenated blood separate)

Differentiation of teeth

Large brain - capable of learning

Diaphragm (efficient breathing)

Endothermic (warm blooded)

Mammary glands (nourish young)

Viviparous

Internal fertilization

3 Subclasses:

1. Monotremes (egg-laying mammals)

platypuses and spiny anteaters

reptilian type egg

hair and mammary glands (no nipples)

2. Marsupials (mammals with pouches)

kangaroos and koalas

reptilian type egg

hair and mammary glands

young craw out of reproductive tract very early in development, craw into pouch and attaches to nipple for nourishment, and develops in pouch)

3. Placental mammals